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1tgo

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(New page: 200px<br /><applet load="1tgo" size="450" color="white" frame="true" align="right" spinBox="true" caption="1tgo, resolution 2.5&Aring;" /> '''THERMOSTABLE B TYPE D...)
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[[Image:1tgo.jpg|left|200px]]<br /><applet load="1tgo" size="350" color="white" frame="true" align="right" spinBox="true"
caption="1tgo, resolution 2.5&Aring;" />
caption="1tgo, resolution 2.5&Aring;" />
'''THERMOSTABLE B TYPE DNA POLYMERASE FROM THERMOCOCCUS GORGONARIUS'''<br />
'''THERMOSTABLE B TYPE DNA POLYMERASE FROM THERMOCOCCUS GORGONARIUS'''<br />
==Overview==
==Overview==
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Most known archaeal DNA polymerases belong to the type B family, which, also includes the DNA replication polymerases of eukaryotes, but maintain, high fidelity at extreme conditions. We describe here the 2.5 A resolution, crystal structure of a DNA polymerase from the Archaea Thermococcus, gorgonarius and identify structural features of the fold and the active, site that are likely responsible for its thermostable function. Comparison, with the mesophilic B type DNA polymerase gp43 of the bacteriophage RB69, highlights thermophilic adaptations, which include the presence of two, disulfide bonds and an enhanced electrostatic complementarity at the, DNA-protein interface. In contrast to gp43, several loops in the, exonuclease and thumb domains are more closely packed; this apparently, blocks primer binding to the exonuclease active site. A physiological role, of this "closed" conformation is unknown but may represent a polymerase, mode, in contrast to an editing mode with an open exonuclease site. This, archaeal B DNA polymerase structure provides a starting point for, structure-based design of polymerases or ligands with applications in, biotechnology and the development of antiviral or anticancer agents.
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Most known archaeal DNA polymerases belong to the type B family, which also includes the DNA replication polymerases of eukaryotes, but maintain high fidelity at extreme conditions. We describe here the 2.5 A resolution crystal structure of a DNA polymerase from the Archaea Thermococcus gorgonarius and identify structural features of the fold and the active site that are likely responsible for its thermostable function. Comparison with the mesophilic B type DNA polymerase gp43 of the bacteriophage RB69 highlights thermophilic adaptations, which include the presence of two disulfide bonds and an enhanced electrostatic complementarity at the DNA-protein interface. In contrast to gp43, several loops in the exonuclease and thumb domains are more closely packed; this apparently blocks primer binding to the exonuclease active site. A physiological role of this "closed" conformation is unknown but may represent a polymerase mode, in contrast to an editing mode with an open exonuclease site. This archaeal B DNA polymerase structure provides a starting point for structure-based design of polymerases or ligands with applications in biotechnology and the development of antiviral or anticancer agents.
==About this Structure==
==About this Structure==
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1TGO is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Thermococcus_gorgonarius Thermococcus gorgonarius]. Active as [http://en.wikipedia.org/wiki/DNA-directed_DNA_polymerase DNA-directed DNA polymerase], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=2.7.7.7 2.7.7.7] Full crystallographic information is available from [http://ispc.weizmann.ac.il/oca-bin/ocashort?id=1TGO OCA].
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1TGO is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Thermococcus_gorgonarius Thermococcus gorgonarius]. Active as [http://en.wikipedia.org/wiki/DNA-directed_DNA_polymerase DNA-directed DNA polymerase], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=2.7.7.7 2.7.7.7] Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1TGO OCA].
==Reference==
==Reference==
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[[Category: Ankenbauer, W.]]
[[Category: Ankenbauer, W.]]
[[Category: Eichinger, A.]]
[[Category: Eichinger, A.]]
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[[Category: Engh, R.A.]]
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[[Category: Engh, R A.]]
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[[Category: Hopfner, K.P.]]
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[[Category: Hopfner, K P.]]
[[Category: Huber, R.]]
[[Category: Huber, R.]]
[[Category: Laue, F.]]
[[Category: Laue, F.]]
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[[Category: replication]]
[[Category: replication]]
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''Page seeded by [http://ispc.weizmann.ac.il/oca OCA ] on Wed Nov 21 03:15:00 2007''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 15:13:20 2008''

Revision as of 13:13, 21 February 2008


1tgo, resolution 2.5Å

Drag the structure with the mouse to rotate

THERMOSTABLE B TYPE DNA POLYMERASE FROM THERMOCOCCUS GORGONARIUS

Overview

Most known archaeal DNA polymerases belong to the type B family, which also includes the DNA replication polymerases of eukaryotes, but maintain high fidelity at extreme conditions. We describe here the 2.5 A resolution crystal structure of a DNA polymerase from the Archaea Thermococcus gorgonarius and identify structural features of the fold and the active site that are likely responsible for its thermostable function. Comparison with the mesophilic B type DNA polymerase gp43 of the bacteriophage RB69 highlights thermophilic adaptations, which include the presence of two disulfide bonds and an enhanced electrostatic complementarity at the DNA-protein interface. In contrast to gp43, several loops in the exonuclease and thumb domains are more closely packed; this apparently blocks primer binding to the exonuclease active site. A physiological role of this "closed" conformation is unknown but may represent a polymerase mode, in contrast to an editing mode with an open exonuclease site. This archaeal B DNA polymerase structure provides a starting point for structure-based design of polymerases or ligands with applications in biotechnology and the development of antiviral or anticancer agents.

About this Structure

1TGO is a Single protein structure of sequence from Thermococcus gorgonarius. Active as DNA-directed DNA polymerase, with EC number 2.7.7.7 Full crystallographic information is available from OCA.

Reference

Crystal structure of a thermostable type B DNA polymerase from Thermococcus gorgonarius., Hopfner KP, Eichinger A, Engh RA, Laue F, Ankenbauer W, Huber R, Angerer B, Proc Natl Acad Sci U S A. 1999 Mar 30;96(7):3600-5. PMID:10097083

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