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1dx5

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[[Image:1dx5.gif|left|200px]]<br /><applet load="1dx5" size="350" color="white" frame="true" align="right" spinBox="true"
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[[Image:1dx5.gif|left|200px]]
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caption="1dx5, resolution 2.30&Aring;" />
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'''CRYSTAL STRUCTURE OF THE THROMBIN-THROMBOMODULIN COMPLEX'''<br />
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{{Structure
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|PDB= 1dx5 |SIZE=350|CAPTION= <scene name='initialview01'>1dx5</scene>, resolution 2.30&Aring;
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|SITE= <scene name='pdbsite=AC1:Catalytic+Triad'>AC1</scene>, <scene name='pdbsite=AC2:Catalytic+Triad'>AC2</scene>, <scene name='pdbsite=AC3:Catalytic+Triad'>AC3</scene> and <scene name='pdbsite=AC4:Catalytic+Triad'>AC4</scene>
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|LIGAND= <scene name='pdbligand=NDG:2-(ACETYLAMINO)-2-DEOXY-A-D-GLUCOPYRANOSE'>NDG</scene>, <scene name='pdbligand=CA:CALCIUM+ION'>CA</scene>, <scene name='pdbligand=NA:SODIUM+ION'>NA</scene> and <scene name='pdbligand=FMT:FORMIC ACID'>FMT</scene>
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|ACTIVITY= [http://en.wikipedia.org/wiki/Thrombin Thrombin], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=3.4.21.5 3.4.21.5]
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|GENE=
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}}
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'''CRYSTAL STRUCTURE OF THE THROMBIN-THROMBOMODULIN COMPLEX'''
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==Overview==
==Overview==
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==About this Structure==
==About this Structure==
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1DX5 is a [http://en.wikipedia.org/wiki/Protein_complex Protein complex] structure of sequences from [http://en.wikipedia.org/wiki/Homo_sapiens Homo sapiens] with <scene name='pdbligand=NDG:'>NDG</scene>, <scene name='pdbligand=CA:'>CA</scene>, <scene name='pdbligand=NA:'>NA</scene> and <scene name='pdbligand=FMT:'>FMT</scene> as [http://en.wikipedia.org/wiki/ligands ligands]. Active as [http://en.wikipedia.org/wiki/Thrombin Thrombin], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=3.4.21.5 3.4.21.5] Known structural/functional Sites: <scene name='pdbsite=AC1:Catalytic+Triad'>AC1</scene>, <scene name='pdbsite=AC2:Catalytic+Triad'>AC2</scene>, <scene name='pdbsite=AC3:Catalytic+Triad'>AC3</scene> and <scene name='pdbsite=AC4:Catalytic+Triad'>AC4</scene>. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1DX5 OCA].
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1DX5 is a [[Protein complex]] structure of sequences from [http://en.wikipedia.org/wiki/Homo_sapiens Homo sapiens]. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1DX5 OCA].
==Reference==
==Reference==
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Structural basis for the anticoagulant activity of the thrombin-thrombomodulin complex., Fuentes-Prior P, Iwanaga Y, Huber R, Pagila R, Rumennik G, Seto M, Morser J, Light DR, Bode W, Nature. 2000 Mar 30;404(6777):518-25. PMID:[http://ispc.weizmann.ac.il//pmbin/getpm?pmid=10761923 10761923]
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Structural basis for the anticoagulant activity of the thrombin-thrombomodulin complex., Fuentes-Prior P, Iwanaga Y, Huber R, Pagila R, Rumennik G, Seto M, Morser J, Light DR, Bode W, Nature. 2000 Mar 30;404(6777):518-25. PMID:[http://www.ncbi.nlm.nih.gov/pubmed/10761923 10761923]
[[Category: Homo sapiens]]
[[Category: Homo sapiens]]
[[Category: Protein complex]]
[[Category: Protein complex]]
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[[Category: anticoagulant complex]]
[[Category: anticoagulant complex]]
[[Category: antifibrinolytic complex]]
[[Category: antifibrinolytic complex]]
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[[Category: egf-like domains]]
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[[Category: egf-like domain]]
[[Category: serine proteinase]]
[[Category: serine proteinase]]
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 12:21:25 2008''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Mar 20 10:45:50 2008''

Revision as of 08:45, 20 March 2008


PDB ID 1dx5

Drag the structure with the mouse to rotate
, resolution 2.30Å
Sites: , , and
Ligands: , , and
Activity: Thrombin, with EC number 3.4.21.5
Coordinates: save as pdb, mmCIF, xml



CRYSTAL STRUCTURE OF THE THROMBIN-THROMBOMODULIN COMPLEX


Contents

Overview

The serine proteinase alpha-thrombin causes blood clotting through proteolytic cleavage of fibrinogen and protease-activated receptors and amplifies its own generation by activating the essential clotting factors V and VIII. Thrombomodulin, a transmembrane thrombin receptor with six contiguous epidermal growth factor-like domains (TME1-6), profoundly alters the substrate specificity of thrombin from pro- to anticoagulant by activating protein C. Activated protein C then deactivates the coagulation cascade by degrading activated factors V and VIII. The thrombin-thrombomodulin complex inhibits fibrinolysis by activating the procarboxypeptidase thrombin-activatable fibrinolysis inhibitor. Here we present the 2.3 A crystal structure of human alpha-thrombin bound to the smallest thrombomodulin fragment required for full protein-C co-factor activity, TME456. The Y-shaped thrombomodulin fragment binds to thrombin's anion-binding exosite-I, preventing binding of procoagulant substrates. Thrombomodulin binding does not seem to induce marked allosteric structural rearrangements at the thrombin active site. Rather, docking of a protein C model to thrombin-TME456 indicates that TME45 may bind substrates in such a manner that their zymogen-activation cleavage sites are presented optimally to the unaltered thrombin active site.

Disease

Known diseases associated with this structure: Dysprothrombinemia OMIM:[176930], Hyperprothrombinemia OMIM:[176930], Hypoprothrombinemia OMIM:[176930], Myocardial infarction, susceptibility to OMIM:[188040], Thrombophilia due to thrombomodulin defect OMIM:[188040]

About this Structure

1DX5 is a Protein complex structure of sequences from Homo sapiens. Full crystallographic information is available from OCA.

Reference

Structural basis for the anticoagulant activity of the thrombin-thrombomodulin complex., Fuentes-Prior P, Iwanaga Y, Huber R, Pagila R, Rumennik G, Seto M, Morser J, Light DR, Bode W, Nature. 2000 Mar 30;404(6777):518-25. PMID:10761923

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