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1m4z

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{{Structure
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|GENE= Orc1 ([http://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&srchmode=5&id=4932 Saccharomyces cerevisiae])
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|RESOURCES=<span class='plainlinks'>[http://oca.weizmann.ac.il/oca-docs/fgij/fg.htm?mol=1m4z FirstGlance], [http://oca.weizmann.ac.il/oca-bin/ocaids?id=1m4z OCA], [http://www.ebi.ac.uk/pdbsum/1m4z PDBsum], [http://www.rcsb.org/pdb/explore.do?structureId=1m4z RCSB]</span>
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'''Crystal structure of the N-terminal BAH domain of Orc1p'''
'''Crystal structure of the N-terminal BAH domain of Orc1p'''
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[[Category: Xu, R M.]]
[[Category: Xu, R M.]]
[[Category: Zhang, Z.]]
[[Category: Zhang, Z.]]
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[[Category: bah domain]]
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[[Category: Bah domain]]
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[[Category: chromatin]]
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[[Category: Chromatin]]
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[[Category: dna replication]]
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[[Category: Dna replication]]
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[[Category: transcriptional silencing]]
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[[Category: Transcriptional silencing]]
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Sat May 3 00:38:14 2008''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Sun Mar 30 22:11:20 2008''
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Revision as of 21:38, 2 May 2008

Template:STRUCTURE 1m4z

Crystal structure of the N-terminal BAH domain of Orc1p


Overview

The N-terminal domain of the largest subunit of the Saccharomyces cerevisiae origin recognition complex (Orc1p) functions in transcriptional silencing and contains a bromo-adjacent homology (BAH) domain found in some chromatin-associated proteins including Sir3p. The 2.2 A crystal structure of the N-terminal domain of Orc1p revealed a BAH core and a non-conserved helical sub-domain. Mutational analyses demonstrated that the helical sub-domain was necessary and sufficient to bind Sir1p, and critical for targeting Sir1p primarily to the cis-acting E silencers at the HMR and HML silent chromatin domains. In the absence of the BAH domain, approximately 14-20% of cells in a population were silenced at the HML locus. Moreover, the distributions of the Sir2p, Sir3p and Sir4p proteins, while normal, were at levels lower than found in wild-type cells. Thus, in the absence of the Orc1p BAH domain, HML resembled silencing of genes adjacent to telomeres. These data are consistent with the view that the Orc1p-Sir1p interaction at the E silencers ensures stable inheritance of pre-established Sir2p, Sir3p and Sir4p complexes at the silent mating type loci.

About this Structure

1M4Z is a Single protein structure of sequence from Saccharomyces cerevisiae. Full crystallographic information is available from OCA.

Reference

Structure and function of the BAH-containing domain of Orc1p in epigenetic silencing., Zhang Z, Hayashi MK, Merkel O, Stillman B, Xu RM, EMBO J. 2002 Sep 2;21(17):4600-11. PMID:12198162 Page seeded by OCA on Sat May 3 00:38:14 2008

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