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5bp3

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==Dehydratase domain (DH) of the mycocerosic acid synthase (MAS), crystal form 2==
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==Dehydratase domain (DH) of a mycocerosic acid synthase-like (MAS-like) PKS, crystal form 2==
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<StructureSection load='5bp3' size='340' side='right' caption='[[5bp3]], [[Resolution|resolution]] 1.45&Aring;' scene=''>
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<StructureSection load='5bp3' size='340' side='right'caption='[[5bp3]], [[Resolution|resolution]] 1.45&Aring;' scene=''>
== Structural highlights ==
== Structural highlights ==
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<table><tr><td colspan='2'>[[5bp3]] is a 2 chain structure. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=5BP3 OCA]. For a <b>guided tour on the structure components</b> use [http://oca.weizmann.ac.il/oca-docs/fgij/fg.htm?mol=5BP3 FirstGlance]. <br>
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<table><tr><td colspan='2'>[[5bp3]] is a 2 chain structure with sequence from [https://en.wikipedia.org/wiki/Mycolicibacterium_smegmatis_MC2_155 Mycolicibacterium smegmatis MC2 155]. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=5BP3 OCA]. For a <b>guided tour on the structure components</b> use [https://proteopedia.org/fgij/fg.htm?mol=5BP3 FirstGlance]. <br>
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</td></tr><tr id='ligand'><td class="sblockLbl"><b>[[Ligand|Ligands:]]</b></td><td class="sblockDat"><scene name='pdbligand=EDO:1,2-ETHANEDIOL'>EDO</scene></td></tr>
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</td></tr><tr id='method'><td class="sblockLbl"><b>[[Empirical_models|Method:]]</b></td><td class="sblockDat" id="methodDat">X-ray diffraction, [[Resolution|Resolution]] 1.45&#8491;</td></tr>
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<tr id='related'><td class="sblockLbl"><b>[[Related_structure|Related:]]</b></td><td class="sblockDat">[[5bp4|5bp4]], [[5bp5|5bp5]], [[5bp2|5bp2]]</td></tr>
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<tr id='ligand'><td class="sblockLbl"><b>[[Ligand|Ligands:]]</b></td><td class="sblockDat" id="ligandDat"><scene name='pdbligand=EDO:1,2-ETHANEDIOL'>EDO</scene></td></tr>
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<tr id='resources'><td class="sblockLbl"><b>Resources:</b></td><td class="sblockDat"><span class='plainlinks'>[http://oca.weizmann.ac.il/oca-docs/fgij/fg.htm?mol=5bp3 FirstGlance], [http://oca.weizmann.ac.il/oca-bin/ocaids?id=5bp3 OCA], [http://pdbe.org/5bp3 PDBe], [http://www.rcsb.org/pdb/explore.do?structureId=5bp3 RCSB], [http://www.ebi.ac.uk/pdbsum/5bp3 PDBsum]</span></td></tr>
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<tr id='resources'><td class="sblockLbl"><b>Resources:</b></td><td class="sblockDat"><span class='plainlinks'>[https://proteopedia.org/fgij/fg.htm?mol=5bp3 FirstGlance], [http://oca.weizmann.ac.il/oca-bin/ocaids?id=5bp3 OCA], [https://pdbe.org/5bp3 PDBe], [https://www.rcsb.org/pdb/explore.do?structureId=5bp3 RCSB], [https://www.ebi.ac.uk/pdbsum/5bp3 PDBsum], [https://prosat.h-its.org/prosat/prosatexe?pdbcode=5bp3 ProSAT]</span></td></tr>
</table>
</table>
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== Function ==
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[https://www.uniprot.org/uniprot/PKS5_MYCS2 PKS5_MYCS2] Polyketide synthase involved in the biosynthesis of 2,4-dimethyl-2-eicosenoic acid, a lipid component of the lipooligosaccharides (LOS) which are not located at the bacterial surface but rather in deeper compartments of the cell envelope of M.smegmatis.<ref>PMID:19181796</ref>
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<div style="background-color:#fffaf0;">
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== Publication Abstract from PubMed ==
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Polyketide synthases (PKSs) are biosynthetic factories that produce natural products with important biological and pharmacological activities. Their exceptional product diversity is encoded in a modular architecture. Modular PKSs (modPKSs) catalyse reactions colinear to the order of modules in an assembly line, whereas iterative PKSs (iPKSs) use a single module iteratively as exemplified by fungal iPKSs (fiPKSs). However, in some cases non-colinear iterative action is also observed for modPKSs modules and is controlled by the assembly line environment. PKSs feature a structural and functional separation into a condensing and a modifying region as observed for fatty acid synthases. Despite the outstanding relevance of PKSs, the detailed organization of PKSs with complete fully reducing modifying regions remains elusive. Here we report a hybrid crystal structure of Mycobacterium smegmatis mycocerosic acid synthase based on structures of its condensing and modifying regions. Mycocerosic acid synthase is a fully reducing iPKS, closely related to modPKSs, and the prototype of mycobacterial mycocerosic acid synthase-like PKSs. It is involved in the biosynthesis of C20-C28 branched-chain fatty acids, which are important virulence factors of mycobacteria. Our structural data reveal a dimeric linker-based organization of the modifying region and visualize dynamics and conformational coupling in PKSs. On the basis of comparative small-angle X-ray scattering, the observed modifying region architecture may be common also in modPKSs. The linker-based organization provides a rationale for the characteristic variability of PKS modules as a main contributor to product diversity. The comprehensive architectural model enables functional dissection and re-engineering of PKSs.
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Mycocerosic acid synthase exemplifies the architecture of reducing polyketide synthases.,Herbst DA, Jakob RP, Zahringer F, Maier T Nature. 2016 Mar 24;531(7595):533-7. doi: 10.1038/nature16993. Epub 2016 Mar 14. PMID:26976449<ref>PMID:26976449</ref>
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From MEDLINE&reg;/PubMed&reg;, a database of the U.S. National Library of Medicine.<br>
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</div>
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<div class="pdbe-citations 5bp3" style="background-color:#fffaf0;"></div>
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== References ==
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<references/>
__TOC__
__TOC__
</StructureSection>
</StructureSection>
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[[Category: Herbst, D A]]
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[[Category: Large Structures]]
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[[Category: Jakob, P R]]
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[[Category: Mycolicibacterium smegmatis MC2 155]]
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[[Category: Maier, T]]
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[[Category: Herbst DA]]
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[[Category: Zaehringer, F]]
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[[Category: Jakob PR]]
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[[Category: Dehydratase]]
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[[Category: Maier T]]
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[[Category: Dh]]
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[[Category: Zaehringer F]]
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[[Category: Lyase]]
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[[Category: Pk]]
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[[Category: Polyketide]]
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Current revision

Dehydratase domain (DH) of a mycocerosic acid synthase-like (MAS-like) PKS, crystal form 2

PDB ID 5bp3

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