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1gpj

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==Overview==
==Overview==
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Processes vital to life such as respiration and photosynthesis critically, depend on the availability of tetrapyrroles including hemes and, chlorophylls. tRNA-dependent catalysis generally is associated with, protein biosynthesis. An exception is the reduction of glutamyl-tRNA to, glutamate-1-semialdehyde by the enzyme glutamyl-tRNA reductase. This, reaction is the indispensable initiating step of tetrapyrrole biosynthesis, in plants and most prokaryotes. The crystal structure of glutamyl-tRNA, reductase from the archaeon Methanopyrus kandleri in complex with the, substrate-like inhibitor glutamycin at 1.9 A resolution reveals an, extended yet planar V-shaped dimer. The well defined interactions of the, inhibitor with the active site support a thioester-mediated reduction, process. Modeling the glutamyl-tRNA onto each monomer reveals an extensive, protein-tRNA interface. We furthermore propose a model whereby the large, void of glutamyl-tRNA reductase is occupied by, glutamate-1-semialdehyde-1,2-mutase, the subsequent enzyme of this, pathway, allowing for the efficient synthesis of 5-aminolevulinic acid, the common precursor of all tetrapyrroles.
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Processes vital to life such as respiration and photosynthesis critically depend on the availability of tetrapyrroles including hemes and chlorophylls. tRNA-dependent catalysis generally is associated with protein biosynthesis. An exception is the reduction of glutamyl-tRNA to glutamate-1-semialdehyde by the enzyme glutamyl-tRNA reductase. This reaction is the indispensable initiating step of tetrapyrrole biosynthesis in plants and most prokaryotes. The crystal structure of glutamyl-tRNA reductase from the archaeon Methanopyrus kandleri in complex with the substrate-like inhibitor glutamycin at 1.9 A resolution reveals an extended yet planar V-shaped dimer. The well defined interactions of the inhibitor with the active site support a thioester-mediated reduction process. Modeling the glutamyl-tRNA onto each monomer reveals an extensive protein-tRNA interface. We furthermore propose a model whereby the large void of glutamyl-tRNA reductase is occupied by glutamate-1-semialdehyde-1,2-mutase, the subsequent enzyme of this pathway, allowing for the efficient synthesis of 5-aminolevulinic acid, the common precursor of all tetrapyrroles.
==About this Structure==
==About this Structure==
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[[Category: Beier, V.]]
[[Category: Beier, V.]]
[[Category: Bringemeier, I.]]
[[Category: Bringemeier, I.]]
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[[Category: Heinz, D.W.]]
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[[Category: Heinz, D W.]]
[[Category: Jahn, D.]]
[[Category: Jahn, D.]]
[[Category: Moser, J.]]
[[Category: Moser, J.]]
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[[Category: Schubert, W.D.]]
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[[Category: Schubert, W D.]]
[[Category: CIT]]
[[Category: CIT]]
[[Category: GMC]]
[[Category: GMC]]
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[[Category: trna-dependent tetrapyrrole biosynthesis]]
[[Category: trna-dependent tetrapyrrole biosynthesis]]
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Sun Feb 3 09:41:36 2008''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 12:52:36 2008''

Revision as of 10:52, 21 February 2008


1gpj, resolution 1.95Å

Drag the structure with the mouse to rotate

GLUTAMYL-TRNA REDUCTASE FROM METHANOPYRUS KANDLERI

Overview

Processes vital to life such as respiration and photosynthesis critically depend on the availability of tetrapyrroles including hemes and chlorophylls. tRNA-dependent catalysis generally is associated with protein biosynthesis. An exception is the reduction of glutamyl-tRNA to glutamate-1-semialdehyde by the enzyme glutamyl-tRNA reductase. This reaction is the indispensable initiating step of tetrapyrrole biosynthesis in plants and most prokaryotes. The crystal structure of glutamyl-tRNA reductase from the archaeon Methanopyrus kandleri in complex with the substrate-like inhibitor glutamycin at 1.9 A resolution reveals an extended yet planar V-shaped dimer. The well defined interactions of the inhibitor with the active site support a thioester-mediated reduction process. Modeling the glutamyl-tRNA onto each monomer reveals an extensive protein-tRNA interface. We furthermore propose a model whereby the large void of glutamyl-tRNA reductase is occupied by glutamate-1-semialdehyde-1,2-mutase, the subsequent enzyme of this pathway, allowing for the efficient synthesis of 5-aminolevulinic acid, the common precursor of all tetrapyrroles.

About this Structure

1GPJ is a Single protein structure of sequence from Methanopyrus kandleri with and as ligands. Known structural/functional Site: . Full crystallographic information is available from OCA.

Reference

V-shaped structure of glutamyl-tRNA reductase, the first enzyme of tRNA-dependent tetrapyrrole biosynthesis., Moser J, Schubert WD, Beier V, Bringemeier I, Jahn D, Heinz DW, EMBO J. 2001 Dec 3;20(23):6583-90. PMID:11726494

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