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1hjf

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[[Image:1hjf.gif|left|200px]]<br /><applet load="1hjf" size="350" color="white" frame="true" align="right" spinBox="true"
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[[Image:1hjf.gif|left|200px]]
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caption="1hjf, resolution 1.60&Aring;" />
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'''ALTERATION OF THE CO-SUBSTRATE SELECTIVITY OF DEACETOXYCEPHALOSPORIN C SYNTHASE: THE ROLE OF ARGININE-258'''<br />
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{{Structure
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|PDB= 1hjf |SIZE=350|CAPTION= <scene name='initialview01'>1hjf</scene>, resolution 1.60&Aring;
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|SITE= <scene name='pdbsite=COI:Coi+Binding+Site+For+Chain+A'>COI</scene> and <scene name='pdbsite=FE:Protein+Fe-Binding+Ligands.+2-Oxo-4-Methylpentanoate+Cos+...'>FE</scene>
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|LIGAND= <scene name='pdbligand=FE2:FE+(II)+ION'>FE2</scene> and <scene name='pdbligand=COI:2-OXO-4-METHYLPENTANOIC ACID'>COI</scene>
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|ACTIVITY=
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|GENE= CEFE, R258Q MUTANT ([http://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&srchmode=5&id=1901 Streptomyces clavuligerus])
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}}
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'''ALTERATION OF THE CO-SUBSTRATE SELECTIVITY OF DEACETOXYCEPHALOSPORIN C SYNTHASE: THE ROLE OF ARGININE-258'''
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==Overview==
==Overview==
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==About this Structure==
==About this Structure==
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1HJF is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Streptomyces_clavuligerus Streptomyces clavuligerus] with <scene name='pdbligand=FE2:'>FE2</scene> and <scene name='pdbligand=COI:'>COI</scene> as [http://en.wikipedia.org/wiki/ligands ligands]. Known structural/functional Sites: <scene name='pdbsite=COI:Coi+Binding+Site+For+Chain+A'>COI</scene> and <scene name='pdbsite=FE:Protein+Fe-Binding+Ligands.+2-Oxo-4-Methylpentanoate+Cos+...'>FE</scene>. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1HJF OCA].
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1HJF is a [[Single protein]] structure of sequence from [http://en.wikipedia.org/wiki/Streptomyces_clavuligerus Streptomyces clavuligerus]. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1HJF OCA].
==Reference==
==Reference==
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Alteration of the co-substrate selectivity of deacetoxycephalosporin C synthase. The role of arginine 258., Lee HJ, Lloyd MD, Clifton IJ, Harlos K, Dubus A, Baldwin JE, Frere JM, Schofield CJ, J Biol Chem. 2001 May 25;276(21):18290-5. Epub 2001 Feb 21. PMID:[http://ispc.weizmann.ac.il//pmbin/getpm?pmid=11279000 11279000]
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Alteration of the co-substrate selectivity of deacetoxycephalosporin C synthase. The role of arginine 258., Lee HJ, Lloyd MD, Clifton IJ, Harlos K, Dubus A, Baldwin JE, Frere JM, Schofield CJ, J Biol Chem. 2001 May 25;276(21):18290-5. Epub 2001 Feb 21. PMID:[http://www.ncbi.nlm.nih.gov/pubmed/11279000 11279000]
[[Category: Single protein]]
[[Category: Single protein]]
[[Category: Streptomyces clavuligerus]]
[[Category: Streptomyces clavuligerus]]
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[[Category: FE2]]
[[Category: FE2]]
[[Category: 2-oxoglutarate-dependent oxygenase]]
[[Category: 2-oxoglutarate-dependent oxygenase]]
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[[Category: alternative 2-oxoacids]]
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[[Category: alternative 2-oxoacid]]
[[Category: cephem antibiotic biosynthesis]]
[[Category: cephem antibiotic biosynthesis]]
[[Category: chemical cosubstrate rescue]]
[[Category: chemical cosubstrate rescue]]
[[Category: co-substrate selectivity]]
[[Category: co-substrate selectivity]]
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 13:01:55 2008''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Mar 20 11:37:50 2008''

Revision as of 09:37, 20 March 2008


PDB ID 1hjf

Drag the structure with the mouse to rotate
, resolution 1.60Å
Sites: and
Ligands: and
Gene: CEFE, R258Q MUTANT (Streptomyces clavuligerus)
Coordinates: save as pdb, mmCIF, xml



ALTERATION OF THE CO-SUBSTRATE SELECTIVITY OF DEACETOXYCEPHALOSPORIN C SYNTHASE: THE ROLE OF ARGININE-258


Overview

Deacetoxycephalosporin C synthase is an iron(II) 2-oxoglutaratedependent oxygenase that catalyzes the oxidative ring-expansion of penicillin N to deacetoxycephalosporin C. The wild-type enzyme is only able to efficiently utilize 2-oxoglutarate and 2-oxoadipate as a 2-oxoacid co-substrate. Mutation of arginine 258, the side chain of which forms an electrostatic interaction with the 5-carboxylate of the 2-oxoglutarate co-substrate, to a glutamine residue reduced activity to about 5% of the wild-type enzyme with 2-oxoglutarate. However, other aliphatic 2-oxoacids, which were not co-substrates for the wild-type enzyme, were utilized by the R258Q mutant. These 2-oxoacids "rescued" catalytic activity to the level observed for the wild-type enzyme as judged by penicillin N and G conversion. These co-substrates underwent oxidative decarboxylation as observed for 2-oxoglutarate in the normal reaction with the wild-type enzyme. Crystal structures of the iron(II)- 2-oxo-3-methylbutanoate (1.5 A), and iron(II)-2-oxo-4-methylpentanoate (1.6 A) enzyme complexes were obtained, which reveal the molecular basis for this "chemical co-substrate rescue" and help to rationalize the co-substrate selectivity of 2-oxoglutaratedependent oxygenases.

About this Structure

1HJF is a Single protein structure of sequence from Streptomyces clavuligerus. Full crystallographic information is available from OCA.

Reference

Alteration of the co-substrate selectivity of deacetoxycephalosporin C synthase. The role of arginine 258., Lee HJ, Lloyd MD, Clifton IJ, Harlos K, Dubus A, Baldwin JE, Frere JM, Schofield CJ, J Biol Chem. 2001 May 25;276(21):18290-5. Epub 2001 Feb 21. PMID:11279000

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