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1qyp

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|PDB= 1qyp |SIZE=350|CAPTION= <scene name='initialview01'>1qyp</scene>
|PDB= 1qyp |SIZE=350|CAPTION= <scene name='initialview01'>1qyp</scene>
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|LIGAND= <scene name='pdbligand=ZN:ZINC ION'>ZN</scene>
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|LIGAND= <scene name='pdbligand=ZN:ZINC+ION'>ZN</scene>
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|ACTIVITY=
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|RELATEDENTRY=
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|RESOURCES=<span class='plainlinks'>[http://oca.weizmann.ac.il/oca-docs/fgij/fg.htm?mol=1qyp FirstGlance], [http://oca.weizmann.ac.il/oca-bin/ocaids?id=1qyp OCA], [http://www.ebi.ac.uk/pdbsum/1qyp PDBsum], [http://www.rcsb.org/pdb/explore.do?structureId=1qyp RCSB]</span>
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[[Category: Wang, B.]]
[[Category: Wang, B.]]
[[Category: Weiss, M A.]]
[[Category: Weiss, M A.]]
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[[Category: ZN]]
 
[[Category: extremophile]]
[[Category: extremophile]]
[[Category: hyperthermophilic]]
[[Category: hyperthermophilic]]
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[[Category: zn ribbon]]
[[Category: zn ribbon]]
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Mar 20 13:44:32 2008''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Sun Mar 30 23:20:49 2008''

Revision as of 20:20, 30 March 2008


PDB ID 1qyp

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Ligands:
Resources: FirstGlance, OCA, PDBsum, RCSB
Coordinates: save as pdb, mmCIF, xml



THERMOCOCCUS CELER RPB9, NMR, 25 STRUCTURES


Overview

BACKGROUND: Transcriptional initiation and elongation provide control points in gene expression. Eukaryotic RNA polymerase II subunit 9 (RPB9) regulates start-site selection and elongational arrest. RPB9 contains Cys4 Zn(2+)-binding motifs which are conserved in archaea and homologous to those of the general transcription factors TFIIB and TFIIS. RESULTS: The structure of an RPB9 domain from the hyperthermophilic archaeon Thermococcus celer was determined at high resolution by NMR spectroscopy. The structure consists of an apical tetrahedral Zn(2+)-binding site, central beta sheet and disordered loop. Although the structure lacks a globular hydrophobic core, the two surfaces of the beta sheet each contain well ordered aromatic rings engaged in serial edge-to-face interactions. Basic sidechains are clustered near the Zn(2+)-binding site. The disordered loop contains sidechains conserved in TFIIS, including acidic residues essential for the stimulation of transcriptional elongation. CONCLUSIONS: The planar architecture of the RPB9 zinc ribbon-distinct from that of a conventional globular domain-can accommodate significant differences in the alignment of polar, non-polar and charged sidechains. Such divergence is associated with local and non-local changes in structure. The RPB9 structure is distinguished by a fourth beta strand (extending the central beta sheet) in a well ordered N-terminal segment and also differs from TFIIS (but not TFIIB) in the orientation of its apical Zn(2+)-binding site. Cys4 Zn(2+)-binding sites with distinct patterns of polar, non-polar and charged residues are conserved among unrelated RNAP subunits and predicted to form variant zinc ribbons.

About this Structure

1QYP is a Single protein structure of sequence from Thermococcus celer. Full crystallographic information is available from OCA.

Reference

High-resolution structure of an archaeal zinc ribbon defines a general architectural motif in eukaryotic RNA polymerases., Wang B, Jones DN, Kaine BP, Weiss MA, Structure. 1998 May 15;6(5):555-69. PMID:9634694

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