4u29

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'''Unreleased structure'''
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==Crystal structure of Rad4-Rad23 crosslinked to a normal DNA duplex==
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<StructureSection load='4u29' size='340' side='right' caption='[[4u29]], [[Resolution|resolution]] 3.05&Aring;' scene=''>
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== Structural highlights ==
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<table><tr><td colspan='2'>[[4u29]] is a 4 chain structure. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=4U29 OCA]. For a <b>guided tour on the structure components</b> use [http://oca.weizmann.ac.il/oca-docs/fgij/fg.htm?mol=4U29 FirstGlance]. <br>
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</td></tr><tr id='NonStdRes'><td class="sblockLbl"><b>[[Non-Standard_Residue|NonStd Res:]]</b></td><td class="sblockDat"><scene name='pdbligand=G47:N2-ETHANETHIOL-2-DEOXY-GUANOSINE-5-MONOPHOSPHATE'>G47</scene></td></tr>
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<tr id='related'><td class="sblockLbl"><b>[[Related_structure|Related:]]</b></td><td class="sblockDat">[[2qsf|2qsf]], [[2qsg|2qsg]], [[2qsh|2qsh]]</td></tr>
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<tr id='resources'><td class="sblockLbl"><b>Resources:</b></td><td class="sblockDat"><span class='plainlinks'>[http://oca.weizmann.ac.il/oca-docs/fgij/fg.htm?mol=4u29 FirstGlance], [http://oca.weizmann.ac.il/oca-bin/ocaids?id=4u29 OCA], [http://www.rcsb.org/pdb/explore.do?structureId=4u29 RCSB], [http://www.ebi.ac.uk/pdbsum/4u29 PDBsum]</span></td></tr>
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</table>
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== Function ==
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[[http://www.uniprot.org/uniprot/RAD4_YEAST RAD4_YEAST]] Involved in nucleotide excision repair of DNA damaged with UV light, bulky adducts, or cross-linking agents. [[http://www.uniprot.org/uniprot/RAD23_YEAST RAD23_YEAST]] Plays a central role both in proteasomal degradation of misfolded proteins and DNA repair. Central component of a complex required to couple deglycosylation and proteasome-mediated degradation of misfolded proteins in the endoplasmic reticulum that are retrotranslocated in the cytosol. Involved in DNA excision repair. May play a part in DNA damage recognition and/or in altering chromatin structure to allow access by damage-processing enzymes.
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<div style="background-color:#fffaf0;">
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== Publication Abstract from PubMed ==
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The xeroderma pigmentosum C (XPC) complex initiates nucleotide excision repair by recognizing DNA lesions before recruiting downstream factors. How XPC detects structurally diverse lesions embedded within normal DNA is unknown. Here we present a crystal structure that captures the yeast XPC orthologue (Rad4) on a single register of undamaged DNA. The structure shows that a disulphide-tethered Rad4 flips out normal nucleotides and adopts a conformation similar to that seen with damaged DNA. Contrary to many DNA repair enzymes that can directly reject non-target sites as structural misfits, our results suggest that Rad4/XPC uses a kinetic gating mechanism whereby lesion selectivity arises from the kinetic competition between DNA opening and the residence time of Rad4/XPC per site. This mechanism is further supported by measurements of Rad4-induced lesion-opening times using temperature-jump perturbation spectroscopy. Kinetic gating may be a general mechanism used by site-specific DNA-binding proteins to minimize time-consuming interrogations of non-target sites.
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The entry 4u29 is ON HOLD
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Kinetic gating mechanism of DNA damage recognition by Rad4/XPC.,Chen X, Velmurugu Y, Zheng G, Park B, Shim Y, Kim Y, Liu L, Van Houten B, He C, Ansari A, Min JH Nat Commun. 2015 Jan 6;6:5849. doi: 10.1038/ncomms6849. PMID:25562780<ref>PMID:25562780</ref>
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Authors: Min, J.-H., Chen, X., Kim, Y.
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From MEDLINE&reg;/PubMed&reg;, a database of the U.S. National Library of Medicine.<br>
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</div>
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Description: Crystal structure of Rad4-Rad23 crosslinked to a normal DNA duplex
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== References ==
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[[Category: Unreleased Structures]]
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<references/>
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[[Category: Min, J.-H]]
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__TOC__
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</StructureSection>
[[Category: Chen, X]]
[[Category: Chen, X]]
[[Category: Kim, Y]]
[[Category: Kim, Y]]
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[[Category: Min, J H]]
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[[Category: Beta-hairpin]]
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[[Category: Disulfide crosslinking]]
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[[Category: Dna binding protein-dna complex]]
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[[Category: Dna damage repair]]
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[[Category: Nucleotide excision repair]]
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[[Category: Protein-dna complex]]
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[[Category: Protein-dna crosslinking]]
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[[Category: Protein-dna interaction]]
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[[Category: Transglutaminase domain]]
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[[Category: Xeroderma pigmentosum]]

Revision as of 11:26, 4 March 2015

Crystal structure of Rad4-Rad23 crosslinked to a normal DNA duplex

4u29, resolution 3.05Å

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