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Odorant binding protein

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==== BmorPBP Function ====
==== BmorPBP Function ====
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Despite three decades of intensive research, the exact roles of OBP and the mechanism by which the odorant receptor (OR) is activated are still in dispute <ref>DOI: 10.1146/annurev-ento-120811-153635</ref><ref>DOI: 10.1007/s00359-009-0461-4</ref>.
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Despite three decades of intensive research, the exact roles of OBP and the mechanism by which the odorant receptor (OR) is activated are still in dispute <ref>DOI: 10.1146/annurev-ento-120811-153635</ref><ref name="kaissling 2009">DOI: 10.1007/s00359-009-0461-4</ref>.
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Of all, the role of OBP as an odorant carrier is generally accepted.
Of all, the role of OBP as an odorant carrier is generally accepted.
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This theory is supported by few assumptions. First, it has been found that the BmPBP conformation is changing from its [["open form" (A)]] to the [["close form" (B)]] in a pH dependent manner <ref>DOI: 10.1074/jbc.274.43.30950</ref>, and that the two conformations has different ligand affinities.
This theory is supported by few assumptions. First, it has been found that the BmPBP conformation is changing from its [["open form" (A)]] to the [["close form" (B)]] in a pH dependent manner <ref>DOI: 10.1074/jbc.274.43.30950</ref>, and that the two conformations has different ligand affinities.
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Second, the surface of the dendrite is negatively charged <ref>DOI: 10.1016/0040-8166(84)90004-1</ref>, which cause the accumulation of positively charged kations near the membrane surface (20-50 nm), thereby inducing a low pH environment near the dendrite membrane [[kaissling 2009]]
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Second, the surface of the dendrite is negatively charged <ref>DOI: 10.1016/0040-8166(84)90004-1</ref>, which cause the accumulation of positively charged kations near the membrane surface (20-50 nm), thereby inducing a low pH environment near the dendrite membrane <ref name="Kissling 2009" /><ref>

Revision as of 08:53, 8 January 2015

Contents

Introduction

Odorant-binding protein (OBP) are soluble proteins which involve in the processes of odorant detection in the olfactory sensilla.

The first OBP that was identified is Bovine odorant binding protein, that was isolated from a cow's mucus ref. OBP in vertebrates

OBP in insects

In order to explain the structure and function of these fascinating proteins, this page will further focus on a particular OBP - the well investigated Bombyx mori PBP: BmorPBP.

PBP

PBPs are specialized members of the insect odorant-binding protein (OBP) super-family, and they are categorized into three groups by their lengths: the long-chain PBPs (∼160 aa), the medium-chain PBPs (∼120 aa), and the short-chain PBPs (∼110 aa)

Bombyx mori BmorPBP (lets talk about sex..)

Bombykol, a sex pheromone of Bombyx mori, from PubChem
Bombykol, a sex pheromone of Bombyx mori, from PubChem

BmorPBP Function

Despite three decades of intensive research, the exact roles of OBP and the mechanism by which the odorant receptor (OR) is activated are still in dispute [1][2].

Of all, the role of OBP as an odorant carrier is generally accepted.

A few functions have been suggested for OBP: 1. 2. 3. 4.


PDB ID 1DQE

Drag the structure with the mouse to rotate


Protein conformations
Receptor activation i lepidoptora
  • The pH dependent theory

This theory is supported by few assumptions. First, it has been found that the BmPBP conformation is changing from its "open form" (A) to the "close form" (B) in a pH dependent manner [3], and that the two conformations has different ligand affinities.

Second, the surface of the dendrite is negatively charged [4], which cause the accumulation of positively charged kations near the membrane surface (20-50 nm), thereby inducing a low pH environment near the dendrite membrane [5][6]

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