2nnx

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[[Image:2nnx.gif|left|200px]]
[[Image:2nnx.gif|left|200px]]
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{{Structure
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<!--
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|PDB= 2nnx |SIZE=350|CAPTION= <scene name='initialview01'>2nnx</scene>, resolution 2.30&Aring;
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The line below this paragraph, containing "STRUCTURE_2nnx", creates the "Structure Box" on the page.
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|SITE=
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You may change the PDB parameter (which sets the PDB file loaded into the applet)
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|LIGAND= <scene name='pdbligand=ACE:ACETYL+GROUP'>ACE</scene>, <scene name='pdbligand=SO4:SULFATE+ION'>SO4</scene>, <scene name='pdbligand=ZN:ZINC+ION'>ZN</scene>
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or the SCENE parameter (which sets the initial scene displayed when the page is loaded),
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|ACTIVITY= <span class='plainlinks'>[http://en.wikipedia.org/wiki/Superoxide_dismutase Superoxide dismutase], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=1.15.1.1 1.15.1.1] </span>
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or leave the SCENE parameter empty for the default display.
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|GENE= SOD1 ([http://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&srchmode=5&id=9606 Homo sapiens])
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-->
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|DOMAIN=
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{{STRUCTURE_2nnx| PDB=2nnx | SCENE= }}
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|RELATEDENTRY=
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|RESOURCES=<span class='plainlinks'>[http://oca.weizmann.ac.il/oca-docs/fgij/fg.htm?mol=2nnx FirstGlance], [http://oca.weizmann.ac.il/oca-bin/ocaids?id=2nnx OCA], [http://www.ebi.ac.uk/pdbsum/2nnx PDBsum], [http://www.rcsb.org/pdb/explore.do?structureId=2nnx RCSB]</span>
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}}
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'''Crystal Structure of the H46R, H48Q double mutant of human [Cu-Zn] Superoxide Dismutase'''
'''Crystal Structure of the H46R, H48Q double mutant of human [Cu-Zn] Superoxide Dismutase'''
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[[Category: Hart, P J.]]
[[Category: Hart, P J.]]
[[Category: Schuermann, J P.]]
[[Category: Schuermann, J P.]]
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[[Category: cu-zn superoxide dismutase]]
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[[Category: Cu-zn superoxide dismutase]]
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[[Category: familial amyotrophic lateral sclerosis mutant]]
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[[Category: Familial amyotrophic lateral sclerosis mutant]]
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[[Category: human]]
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[[Category: Human]]
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[[Category: oxidoreductase]]
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[[Category: Oxidoreductase]]
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[[Category: superoxide acceptor]]
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[[Category: Superoxide acceptor]]
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Sun May 4 09:41:13 2008''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Mon Mar 31 04:05:44 2008''
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Revision as of 06:41, 4 May 2008

Template:STRUCTURE 2nnx

Crystal Structure of the H46R, H48Q double mutant of human [Cu-Zn] Superoxide Dismutase


Contents

Overview

A subset of superoxide dismutase 1 (Cu/Zn-SOD1) mutants that cause familial amyotrophic lateral sclerosis (FALS) have heightened reactivity with (-)ONOO and H(2)O(2) in vitro. This reactivity requires a copper ion bound in the active site and is a suggested mechanism of motor neuron injury. However, we have found that transgenic mice that express SOD1-H46R/H48Q, which combines natural FALS mutations at ligands for copper and which is inactive, develop motor neuron disease. Using a direct radioactive copper incorporation assay in transfected cells and the established tools of single crystal x-ray diffraction, we now demonstrate that this variant does not stably bind copper. We find that single mutations at copper ligands, including H46R, H48Q, and a quadruple mutant H46R/H48Q/H63G/H120G, also diminish the binding of radioactive copper. Further, using native polyacrylamide gel electrophoresis and a yeast two-hybrid assay, the binding of copper was found to be related to the formation of the stable dimeric enzyme. Collectively, our data demonstrate a relationship between copper and assembly of SOD1 into stable dimers and also define disease-causing SOD1 mutants that are unlikely to robustly produce toxic radicals via copper-mediated chemistry.

Disease

Known disease associated with this structure: Amyotrophic lateral sclerosis, due to SOD1 deficiency OMIM:[147450]

About this Structure

2NNX is a Single protein structure of sequence from Homo sapiens. Full crystallographic information is available from OCA.

Reference

Disease-associated mutations at copper ligand histidine residues of superoxide dismutase 1 diminish the binding of copper and compromise dimer stability., Wang J, Caruano-Yzermans A, Rodriguez A, Scheurmann JP, Slunt HH, Cao X, Gitlin J, Hart PJ, Borchelt DR, J Biol Chem. 2007 Jan 5;282(1):345-52. Epub 2006 Nov 8. PMID:17092942 Page seeded by OCA on Sun May 4 09:41:13 2008

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