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1hes

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(New page: 200px<br /> <applet load="1hes" size="450" color="white" frame="true" align="right" spinBox="true" caption="1hes, resolution 3.0&Aring;" /> '''MU2 ADAPTIN SUBUNIT ...)
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'''MU2 ADAPTIN SUBUNIT (AP50) OF AP2 ADAPTOR (SECOND DOMAIN), COMPLEXED WITH P-SELECTIN INTERNALIZATION PEPTIDE SHLGTYGVFTNAA'''<br />
'''MU2 ADAPTIN SUBUNIT (AP50) OF AP2 ADAPTOR (SECOND DOMAIN), COMPLEXED WITH P-SELECTIN INTERNALIZATION PEPTIDE SHLGTYGVFTNAA'''<br />
==Overview==
==Overview==
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Internalization signals of the Yxx phi type (phi = bulky hydrophobic side, chain) interact with the mu 2 chain of AP-2 adaptors. Internalization, activity is intolerant of non-conservative substitution of either the, tyrosine or the phi side chains, which bind to hydrophobic pockets in mu 2, adaptin in a conformation described as 'a two pinned plug into a socket'., P-selectin, a type I transmembrane protein, contains the Yxx phi-like, sequence YGVF in its cytoplasmic domain, but substitution of either the, tyrosine or phenylalanine with alanine in the full-length protein causes, only small changes in the rate of endocytosis. It is shown here that the, sequence YGVF contained within a peptide corresponding to the 17, COOH-terminal amino acids of P-selectin binds to mu 2 adaptin in the same, fashion previously seen for other Yxx phi motifs. In addition, the, P-selectin peptide binds to a third hydrophobic pocket in mu 2 adaptin, through a leucine at position Y-3 in the peptide. This structure suggests, that some sequences can function as a 'three pinned plug', in which, internalization activity is not critically dependent on any one of the, three interacting side chains.
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Internalization signals of the Yxx phi type (phi = bulky hydrophobic side chain) interact with the mu 2 chain of AP-2 adaptors. Internalization activity is intolerant of non-conservative substitution of either the tyrosine or the phi side chains, which bind to hydrophobic pockets in mu 2 adaptin in a conformation described as 'a two pinned plug into a socket'. P-selectin, a type I transmembrane protein, contains the Yxx phi-like sequence YGVF in its cytoplasmic domain, but substitution of either the tyrosine or phenylalanine with alanine in the full-length protein causes only small changes in the rate of endocytosis. It is shown here that the sequence YGVF contained within a peptide corresponding to the 17 COOH-terminal amino acids of P-selectin binds to mu 2 adaptin in the same fashion previously seen for other Yxx phi motifs. In addition, the P-selectin peptide binds to a third hydrophobic pocket in mu 2 adaptin through a leucine at position Y-3 in the peptide. This structure suggests that some sequences can function as a 'three pinned plug', in which internalization activity is not critically dependent on any one of the three interacting side chains.
==Disease==
==Disease==
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==About this Structure==
==About this Structure==
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1HES is a [http://en.wikipedia.org/wiki/Protein_complex Protein complex] structure of sequences from [http://en.wikipedia.org/wiki/Homo_sapiens Homo sapiens] and [http://en.wikipedia.org/wiki/Rattus_norvegicus Rattus norvegicus]. Full crystallographic information is available from [http://ispc.weizmann.ac.il/oca-bin/ocashort?id=1HES OCA].
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1HES is a [http://en.wikipedia.org/wiki/Protein_complex Protein complex] structure of sequences from [http://en.wikipedia.org/wiki/Homo_sapiens Homo sapiens] and [http://en.wikipedia.org/wiki/Rattus_norvegicus Rattus norvegicus]. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1HES OCA].
==Reference==
==Reference==
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[[Category: Protein complex]]
[[Category: Protein complex]]
[[Category: Rattus norvegicus]]
[[Category: Rattus norvegicus]]
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[[Category: Evans, P.R.]]
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[[Category: Evans, P R.]]
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[[Category: Green, S.A.]]
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[[Category: Green, S A.]]
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[[Category: Owen, D.J.]]
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[[Category: Owen, D J.]]
[[Category: adaptor]]
[[Category: adaptor]]
[[Category: endocytosis]]
[[Category: endocytosis]]
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[[Category: peptide complex]]
[[Category: peptide complex]]
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''Page seeded by [http://ispc.weizmann.ac.il/oca OCA ] on Mon Nov 12 17:17:09 2007''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 13:00:36 2008''

Revision as of 11:00, 21 February 2008


1hes, resolution 3.0Å

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MU2 ADAPTIN SUBUNIT (AP50) OF AP2 ADAPTOR (SECOND DOMAIN), COMPLEXED WITH P-SELECTIN INTERNALIZATION PEPTIDE SHLGTYGVFTNAA

Contents

Overview

Internalization signals of the Yxx phi type (phi = bulky hydrophobic side chain) interact with the mu 2 chain of AP-2 adaptors. Internalization activity is intolerant of non-conservative substitution of either the tyrosine or the phi side chains, which bind to hydrophobic pockets in mu 2 adaptin in a conformation described as 'a two pinned plug into a socket'. P-selectin, a type I transmembrane protein, contains the Yxx phi-like sequence YGVF in its cytoplasmic domain, but substitution of either the tyrosine or phenylalanine with alanine in the full-length protein causes only small changes in the rate of endocytosis. It is shown here that the sequence YGVF contained within a peptide corresponding to the 17 COOH-terminal amino acids of P-selectin binds to mu 2 adaptin in the same fashion previously seen for other Yxx phi motifs. In addition, the P-selectin peptide binds to a third hydrophobic pocket in mu 2 adaptin through a leucine at position Y-3 in the peptide. This structure suggests that some sequences can function as a 'three pinned plug', in which internalization activity is not critically dependent on any one of the three interacting side chains.

Disease

Known diseases associated with this structure: Atopy, susceptibility to OMIM:[173610], Platelet alpha/delta storage pool deficiency OMIM:[173610]

About this Structure

1HES is a Protein complex structure of sequences from Homo sapiens and Rattus norvegicus. Full crystallographic information is available from OCA.

Reference

A third specificity-determining site in mu 2 adaptin for sequences upstream of Yxx phi sorting motifs., Owen DJ, Setiadi H, Evans PR, McEver RP, Green SA, Traffic. 2001 Feb;2(2):105-10. PMID:11247301

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