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1jqc

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(New page: 200px<br /><applet load="1jqc" size="450" color="white" frame="true" align="right" spinBox="true" caption="1jqc, resolution 1.61&Aring;" /> '''Mn substituted Ribon...)
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[[Image:1jqc.jpg|left|200px]]<br /><applet load="1jqc" size="350" color="white" frame="true" align="right" spinBox="true"
caption="1jqc, resolution 1.61&Aring;" />
caption="1jqc, resolution 1.61&Aring;" />
'''Mn substituted Ribonucleotide reductase R2 from E. Coli oxidized by hydrogen peroxide and hydroxylamine'''<br />
'''Mn substituted Ribonucleotide reductase R2 from E. Coli oxidized by hydrogen peroxide and hydroxylamine'''<br />
==Overview==
==Overview==
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The di-iron carboxylate proteins constitute a diverse class of non-heme, iron enzymes performing a multitude of redox reactions. These reactions, usually involve high-valent Fe-oxo species and are thought to be, controlled by carboxylate shifts. Owing to their short lifetime, the, intermediate structures have so far escaped structural characterization by, X-ray crystallography. In an attempt to map the carboxylate conformations, available to the protein during different redox states and different, ligand environments, we have studied metal-substituted forms of the R2, protein of ribonucleotide reductase from Escherichia coli. In the present, work we have solved the crystal structures of Mn-substituted R2 oxidized, in two different ways. Oxidation was performed using either nitric oxide, or a combination of hydrogen peroxide and hydroxylamine. The two, structures are virtually identical, indicating that the oxidation states, are the same, most likely a mixed-valent MnII-MnIII centre. One of the, carboxylate ligands (D84) adopts a new, so far unseen, conformation, which, could participate in the mechanism for radical generation in R2. E238, adopts a bridging-chelating conformation proposed to be important for, proper O2 activation but not previously observed in the wild-type enzyme., Probable catalase activity was also observed during the oxidation with, H2O2, indicating mechanistic similarities to the di-Mn catalases.
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The di-iron carboxylate proteins constitute a diverse class of non-heme iron enzymes performing a multitude of redox reactions. These reactions usually involve high-valent Fe-oxo species and are thought to be controlled by carboxylate shifts. Owing to their short lifetime, the intermediate structures have so far escaped structural characterization by X-ray crystallography. In an attempt to map the carboxylate conformations available to the protein during different redox states and different ligand environments, we have studied metal-substituted forms of the R2 protein of ribonucleotide reductase from Escherichia coli. In the present work we have solved the crystal structures of Mn-substituted R2 oxidized in two different ways. Oxidation was performed using either nitric oxide or a combination of hydrogen peroxide and hydroxylamine. The two structures are virtually identical, indicating that the oxidation states are the same, most likely a mixed-valent MnII-MnIII centre. One of the carboxylate ligands (D84) adopts a new, so far unseen, conformation, which could participate in the mechanism for radical generation in R2. E238 adopts a bridging-chelating conformation proposed to be important for proper O2 activation but not previously observed in the wild-type enzyme. Probable catalase activity was also observed during the oxidation with H2O2, indicating mechanistic similarities to the di-Mn catalases.
==About this Structure==
==About this Structure==
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1JQC is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Escherichia_coli Escherichia coli] with MN and HG as [http://en.wikipedia.org/wiki/ligands ligands]. Active as [http://en.wikipedia.org/wiki/Ribonucleoside-diphosphate_reductase Ribonucleoside-diphosphate reductase], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=1.17.4.1 1.17.4.1] Full crystallographic information is available from [http://ispc.weizmann.ac.il/oca-bin/ocashort?id=1JQC OCA].
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1JQC is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Escherichia_coli Escherichia coli] with <scene name='pdbligand=MN:'>MN</scene> and <scene name='pdbligand=HG:'>HG</scene> as [http://en.wikipedia.org/wiki/ligands ligands]. Active as [http://en.wikipedia.org/wiki/Ribonucleoside-diphosphate_reductase Ribonucleoside-diphosphate reductase], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=1.17.4.1 1.17.4.1] Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1JQC OCA].
==Reference==
==Reference==
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[[Category: Ribonucleoside-diphosphate reductase]]
[[Category: Ribonucleoside-diphosphate reductase]]
[[Category: Single protein]]
[[Category: Single protein]]
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[[Category: Andersson, M.E.]]
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[[Category: Andersson, M E.]]
[[Category: Hogbom, M.]]
[[Category: Hogbom, M.]]
[[Category: Nordlund, P.]]
[[Category: Nordlund, P.]]
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[[Category: ribonucleotide reductase r2]]
[[Category: ribonucleotide reductase r2]]
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''Page seeded by [http://ispc.weizmann.ac.il/oca OCA ] on Tue Nov 20 18:28:52 2007''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 13:25:34 2008''

Revision as of 11:25, 21 February 2008


1jqc, resolution 1.61Å

Drag the structure with the mouse to rotate

Mn substituted Ribonucleotide reductase R2 from E. Coli oxidized by hydrogen peroxide and hydroxylamine

Overview

The di-iron carboxylate proteins constitute a diverse class of non-heme iron enzymes performing a multitude of redox reactions. These reactions usually involve high-valent Fe-oxo species and are thought to be controlled by carboxylate shifts. Owing to their short lifetime, the intermediate structures have so far escaped structural characterization by X-ray crystallography. In an attempt to map the carboxylate conformations available to the protein during different redox states and different ligand environments, we have studied metal-substituted forms of the R2 protein of ribonucleotide reductase from Escherichia coli. In the present work we have solved the crystal structures of Mn-substituted R2 oxidized in two different ways. Oxidation was performed using either nitric oxide or a combination of hydrogen peroxide and hydroxylamine. The two structures are virtually identical, indicating that the oxidation states are the same, most likely a mixed-valent MnII-MnIII centre. One of the carboxylate ligands (D84) adopts a new, so far unseen, conformation, which could participate in the mechanism for radical generation in R2. E238 adopts a bridging-chelating conformation proposed to be important for proper O2 activation but not previously observed in the wild-type enzyme. Probable catalase activity was also observed during the oxidation with H2O2, indicating mechanistic similarities to the di-Mn catalases.

About this Structure

1JQC is a Single protein structure of sequence from Escherichia coli with and as ligands. Active as Ribonucleoside-diphosphate reductase, with EC number 1.17.4.1 Full crystallographic information is available from OCA.

Reference

Crystal structures of oxidized dinuclear manganese centres in Mn-substituted class I ribonucleotide reductase from Escherichia coli: carboxylate shifts with implications for O2 activation and radical generation., Hogbom M, Andersson ME, Nordlund P, J Biol Inorg Chem. 2001 Mar;6(3):315-23. PMID:11315567

Page seeded by OCA on Thu Feb 21 13:25:34 2008

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