1ynu

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(New page: 200px<br /><applet load="1ynu" size="450" color="white" frame="true" align="right" spinBox="true" caption="1ynu, resolution 2.25&Aring;" /> '''Crystal structure of...)
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[[Image:1ynu.gif|left|200px]]<br /><applet load="1ynu" size="450" color="white" frame="true" align="right" spinBox="true"
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[[Image:1ynu.gif|left|200px]]<br /><applet load="1ynu" size="350" color="white" frame="true" align="right" spinBox="true"
caption="1ynu, resolution 2.25&Aring;" />
caption="1ynu, resolution 2.25&Aring;" />
'''Crystal structure of apple ACC synthase in complex with L-vinylglycine'''<br />
'''Crystal structure of apple ACC synthase in complex with L-vinylglycine'''<br />
==Overview==
==Overview==
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L-Vinylglycine (L-VG) is both a substrate for and a mechanism-based, inhibitor of 1-aminocyclopropane-1-carboxylate (ACC) synthase. The ratio, of the rate constants for catalytic conversion to alpha-ketobutyrate and, ammonia to inactivation is 500/1. The crystal structure of the covalent, adduct of the inactivated enzyme was determined at 2.25 Angstroms, resolution. The active site contains an external aldimine of the adduct of, L-VG with the pyridoxal 5'-phosphate cofactor. The side chain gamma-carbon, of L-VG is covalently bound to the epsilon-amino group of Lys273. This, species corresponds to one of the two alternatives proposed by Feng and, Kirsch [Feng, L. and Kirsch, J.F. (2000) L-Vinylglycine is an alternative, substrate as well as a mechanism-based inhibitor of, 1-aminocyclopropane-1-carboxylate synthase. Biochemistry 39, 2436-2444], and presumably results from Michael addition to a vinylglycine ketimine, intermediate.
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L-Vinylglycine (L-VG) is both a substrate for and a mechanism-based inhibitor of 1-aminocyclopropane-1-carboxylate (ACC) synthase. The ratio of the rate constants for catalytic conversion to alpha-ketobutyrate and ammonia to inactivation is 500/1. The crystal structure of the covalent adduct of the inactivated enzyme was determined at 2.25 Angstroms resolution. The active site contains an external aldimine of the adduct of L-VG with the pyridoxal 5'-phosphate cofactor. The side chain gamma-carbon of L-VG is covalently bound to the epsilon-amino group of Lys273. This species corresponds to one of the two alternatives proposed by Feng and Kirsch [Feng, L. and Kirsch, J.F. (2000) L-Vinylglycine is an alternative substrate as well as a mechanism-based inhibitor of 1-aminocyclopropane-1-carboxylate synthase. Biochemistry 39, 2436-2444] and presumably results from Michael addition to a vinylglycine ketimine intermediate.
==About this Structure==
==About this Structure==
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1YNU is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Malus_x_domestica Malus x domestica] with NI, K, PY4 and TRS as [http://en.wikipedia.org/wiki/ligands ligands]. Active as [http://en.wikipedia.org/wiki/1-aminocyclopropane-1-carboxylate_synthase 1-aminocyclopropane-1-carboxylate synthase], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=4.4.1.14 4.4.1.14] Full crystallographic information is available from [http://ispc.weizmann.ac.il/oca-bin/ocashort?id=1YNU OCA].
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1YNU is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Malus_x_domestica Malus x domestica] with <scene name='pdbligand=NI:'>NI</scene>, <scene name='pdbligand=K:'>K</scene>, <scene name='pdbligand=PY4:'>PY4</scene> and <scene name='pdbligand=TRS:'>TRS</scene> as [http://en.wikipedia.org/wiki/ligands ligands]. Active as [http://en.wikipedia.org/wiki/1-aminocyclopropane-1-carboxylate_synthase 1-aminocyclopropane-1-carboxylate synthase], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=4.4.1.14 4.4.1.14] Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1YNU OCA].
==Reference==
==Reference==
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[[Category: Single protein]]
[[Category: Single protein]]
[[Category: Capitani, G.]]
[[Category: Capitani, G.]]
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[[Category: Eliot, A.C.]]
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[[Category: Eliot, A C.]]
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[[Category: Grutter, M.G.]]
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[[Category: Grutter, M G.]]
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[[Category: Kirsch, J.F.]]
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[[Category: Kirsch, J F.]]
[[Category: Tschopp, M.]]
[[Category: Tschopp, M.]]
[[Category: K]]
[[Category: K]]
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[[Category: lyase]]
[[Category: lyase]]
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''Page seeded by [http://ispc.weizmann.ac.il/oca OCA ] on Wed Nov 21 06:56:17 2007''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 16:07:22 2008''

Revision as of 14:07, 21 February 2008


1ynu, resolution 2.25Å

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Crystal structure of apple ACC synthase in complex with L-vinylglycine

Overview

L-Vinylglycine (L-VG) is both a substrate for and a mechanism-based inhibitor of 1-aminocyclopropane-1-carboxylate (ACC) synthase. The ratio of the rate constants for catalytic conversion to alpha-ketobutyrate and ammonia to inactivation is 500/1. The crystal structure of the covalent adduct of the inactivated enzyme was determined at 2.25 Angstroms resolution. The active site contains an external aldimine of the adduct of L-VG with the pyridoxal 5'-phosphate cofactor. The side chain gamma-carbon of L-VG is covalently bound to the epsilon-amino group of Lys273. This species corresponds to one of the two alternatives proposed by Feng and Kirsch [Feng, L. and Kirsch, J.F. (2000) L-Vinylglycine is an alternative substrate as well as a mechanism-based inhibitor of 1-aminocyclopropane-1-carboxylate synthase. Biochemistry 39, 2436-2444] and presumably results from Michael addition to a vinylglycine ketimine intermediate.

About this Structure

1YNU is a Single protein structure of sequence from Malus x domestica with , , and as ligands. Active as 1-aminocyclopropane-1-carboxylate synthase, with EC number 4.4.1.14 Full crystallographic information is available from OCA.

Reference

Structure of ACC synthase inactivated by the mechanism-based inhibitor L-vinylglycine., Capitani G, Tschopp M, Eliot AC, Kirsch JF, Grutter MG, FEBS Lett. 2005 Apr 25;579(11):2458-62. PMID:15848188

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