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Shiga toxin

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==Structure==
==Structure==
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Shiga Toxin consists consists of an [http://en.wikipedia.org/wiki/AB5_toxin AB5] hexamer, by X-ray crystallography. The five B subunits form a pentameric ring, encircling a helix at the carboxy terminus of the A subunit. The A subunit interacts with the B pentamer via this C-terminal helix and a four-stranded mixed beta-sheet. The fold of the rest of the A subunit is similar to that of the A chain of the plant toxin ricin; both are N-glycosidases. However, the active site in the bacterial holotoxin is blocked by a segment of polypeptide chain. These residues of the A subunit would be released as part of the activation mechanism of the toxin.
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Shiga Toxin consists consists of an [http://en.wikipedia.org/wiki/AB5_toxin AB5] hexamer.<ref name=Fraser>PMID: 7656009</ref> The 5 subunit B pentamer interacts with a single A subunit via a C-terminal helix and 4 stranded beta-sheet.<ref name=Fraser> The glycosidase active site is located on the A subunit, but is blocked by the B subunit until they are cleaved and an active A subunit is released into the target cell.<ref name=Fraser>
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==Function==
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Shiga Toxin acts as an N-glycosidase, removing an adenine from the 60S ribosomal rRNA of a target cell leading to reduced protein synthesis.<ref name=Di>PMID: 21184769</ref> The B subunit is necessary for binding to eukaryotic cell surface, where it is then endocytosed and proteolytically cleaved into two active A subunits and a B subunit. On the A subunit Tyr77, Tyr114, Glu167, Arg170, and Trp203 are all essential in glycosidic activity.<ref name=Di>
==References==
==References==
{{Reflist}}
{{Reflist}}

Revision as of 02:44, 14 October 2011

Shiga Toxin Type 2 (Stx2)

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Introduction

Shiga Toxins are a family of AB5 toxins which cause dysentery in humans. They are primarily secreted by Shiga toxin-encoding Escherichia coli (STEC), notably by the 0157:H7 strain.[1]

Structure

Shiga Toxin consists consists of an AB5 hexamer.[2] The 5 subunit B pentamer interacts with a single A subunit via a C-terminal helix and 4 stranded beta-sheet.[2] The B subunit is necessary for binding to eukaryotic cell surface, where it is then endocytosed and proteolytically cleaved into two active A subunits and a B subunit. On the A subunit Tyr77, Tyr114, Glu167, Arg170, and Trp203 are all essential in glycosidic activity.[3]

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Max Evoy-Mount, Michal Harel, Alexander Berchansky

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