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1iqq

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(New page: 200px<br /><applet load="1iqq" size="450" color="white" frame="true" align="right" spinBox="true" caption="1iqq, resolution 1.50&Aring;" /> '''Crystal Structure of...)
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'''Crystal Structure of Japanese pear S3-RNase'''<br />
'''Crystal Structure of Japanese pear S3-RNase'''<br />
==Overview==
==Overview==
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The crystal structure of the Pyrus pyrifolia pistil ribonuclease, (S(3)-RNase) responsible for gametophytic self-incompatibility was, determined at 1.5-A resolution. It consists of eight helices and seven, beta-strands, and its folding topology is typical of RNase T(2) family, enzymes. Based on a structural comparison of S(3)-RNase with RNase Rh, a, fungal RNase T(2) family enzyme, the active site residues of S(3)-RNase, assigned were His(33) and His(88) as catalysts and Glu(84) and Lys(87) as, stabilizers of an intermediate in the transition state. Moreover, amino, acid residues that constitute substrate binding sites of the two RNases, could be superimposed geometrically. A hypervariable (HV) region that has, an S-allele-specific sequence comprises a long loop and short alpha-helix., This region is far from the active site cleft, exposed on the molecule's, surface, and positively charged. Four positively selected (PS) regions, in, which the number of nonsynonymous substitutions exceeds that of synonymous, ones, are located on either side of the active site cleft, and accessible, to solvent. These structural features suggest that the HV or PS regions, may interact with a pollen S-gene product(s) to recognize self and, non-self pollen.
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The crystal structure of the Pyrus pyrifolia pistil ribonuclease (S(3)-RNase) responsible for gametophytic self-incompatibility was determined at 1.5-A resolution. It consists of eight helices and seven beta-strands, and its folding topology is typical of RNase T(2) family enzymes. Based on a structural comparison of S(3)-RNase with RNase Rh, a fungal RNase T(2) family enzyme, the active site residues of S(3)-RNase assigned were His(33) and His(88) as catalysts and Glu(84) and Lys(87) as stabilizers of an intermediate in the transition state. Moreover, amino acid residues that constitute substrate binding sites of the two RNases could be superimposed geometrically. A hypervariable (HV) region that has an S-allele-specific sequence comprises a long loop and short alpha-helix. This region is far from the active site cleft, exposed on the molecule's surface, and positively charged. Four positively selected (PS) regions, in which the number of nonsynonymous substitutions exceeds that of synonymous ones, are located on either side of the active site cleft, and accessible to solvent. These structural features suggest that the HV or PS regions may interact with a pollen S-gene product(s) to recognize self and non-self pollen.
==About this Structure==
==About this Structure==
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1IQQ is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Pyrus_pyrifolia Pyrus pyrifolia]. Active as [http://en.wikipedia.org/wiki/Ribonuclease_T(2) Ribonuclease T(2)], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=3.1.27.1 3.1.27.1] Full crystallographic information is available from [http://ispc.weizmann.ac.il/oca-bin/ocashort?id=1IQQ OCA].
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1IQQ is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Pyrus_pyrifolia Pyrus pyrifolia]. Active as [http://en.wikipedia.org/wiki/Ribonuclease_T(2) Ribonuclease T(2)], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=3.1.27.1 3.1.27.1] Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1IQQ OCA].
==Reference==
==Reference==
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[[Category: t2 family ribonuclease]]
[[Category: t2 family ribonuclease]]
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''Page seeded by [http://ispc.weizmann.ac.il/oca OCA ] on Tue Nov 20 17:35:09 2007''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 13:14:43 2008''

Revision as of 11:14, 21 February 2008


1iqq, resolution 1.50Å

Drag the structure with the mouse to rotate

Crystal Structure of Japanese pear S3-RNase

Overview

The crystal structure of the Pyrus pyrifolia pistil ribonuclease (S(3)-RNase) responsible for gametophytic self-incompatibility was determined at 1.5-A resolution. It consists of eight helices and seven beta-strands, and its folding topology is typical of RNase T(2) family enzymes. Based on a structural comparison of S(3)-RNase with RNase Rh, a fungal RNase T(2) family enzyme, the active site residues of S(3)-RNase assigned were His(33) and His(88) as catalysts and Glu(84) and Lys(87) as stabilizers of an intermediate in the transition state. Moreover, amino acid residues that constitute substrate binding sites of the two RNases could be superimposed geometrically. A hypervariable (HV) region that has an S-allele-specific sequence comprises a long loop and short alpha-helix. This region is far from the active site cleft, exposed on the molecule's surface, and positively charged. Four positively selected (PS) regions, in which the number of nonsynonymous substitutions exceeds that of synonymous ones, are located on either side of the active site cleft, and accessible to solvent. These structural features suggest that the HV or PS regions may interact with a pollen S-gene product(s) to recognize self and non-self pollen.

About this Structure

1IQQ is a Single protein structure of sequence from Pyrus pyrifolia. Active as Ribonuclease T(2), with EC number 3.1.27.1 Full crystallographic information is available from OCA.

Reference

Crystal structure at 1.5-A resolution of Pyrus pyrifolia pistil ribonuclease responsible for gametophytic self-incompatibility., Matsuura T, Sakai H, Unno M, Ida K, Sato M, Sakiyama F, Norioka S, J Biol Chem. 2001 Nov 30;276(48):45261-9. Epub 2001 Sep 27. PMID:11577107

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