1ncn

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(New page: 200px<br /> <applet load="1ncn" size="450" color="white" frame="true" align="right" spinBox="true" caption="1ncn, resolution 2.70&Aring;" /> '''the receptor-bindin...)
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<applet load="1ncn" size="450" color="white" frame="true" align="right" spinBox="true"
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caption="1ncn, resolution 2.70&Aring;" />
caption="1ncn, resolution 2.70&Aring;" />
'''the receptor-binding domain of human B7-2'''<br />
'''the receptor-binding domain of human B7-2'''<br />
==Overview==
==Overview==
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B7-1 and B7-2 are homologous costimulatory ligands expressed on the, surfaces of antigen-presenting cells. Their interactions with CD28/CTLA-4, receptors expressed on T cell surfaces are crucial for the proper, regulation of T cell activity. B7-1 and B7-2 display distinct roles in, immune regulation, although they are usually considered to have redundant, functions. Here, we report the crystal structure of the receptor-binding, (Ig V-type) domain of human B7-2 at 2.7-A resolution. Structures of, unliganded and liganded B7-1 and B7-2 suggest a physical-chemical basis, for the observed functional similarities and differences between these two, costimulatory ligands. Of particular note, whereas the majority of the, residues mediating B7-1 dimerization are hydrophobic, the B7-2 dimer, observed in the B7-2/CTLA-4 complex displays a very hydrophilic dimer, interface. These differences provide a mechanism for preventing the, formation of B7-1/B7-2 heterodimers. The divergence at the putative dimer, interface is also consistent with the lower tendency of B7-2 to dimerize, as shown by the monomeric state of unliganded B7-2 both in solution and, crystalline form, and may result in detailed differences in signaling, mechanisms associated with B7-1 and B7-2.
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B7-1 and B7-2 are homologous costimulatory ligands expressed on the surfaces of antigen-presenting cells. Their interactions with CD28/CTLA-4 receptors expressed on T cell surfaces are crucial for the proper regulation of T cell activity. B7-1 and B7-2 display distinct roles in immune regulation, although they are usually considered to have redundant functions. Here, we report the crystal structure of the receptor-binding (Ig V-type) domain of human B7-2 at 2.7-A resolution. Structures of unliganded and liganded B7-1 and B7-2 suggest a physical-chemical basis for the observed functional similarities and differences between these two costimulatory ligands. Of particular note, whereas the majority of the residues mediating B7-1 dimerization are hydrophobic, the B7-2 dimer observed in the B7-2/CTLA-4 complex displays a very hydrophilic dimer interface. These differences provide a mechanism for preventing the formation of B7-1/B7-2 heterodimers. The divergence at the putative dimer interface is also consistent with the lower tendency of B7-2 to dimerize, as shown by the monomeric state of unliganded B7-2 both in solution and crystalline form, and may result in detailed differences in signaling mechanisms associated with B7-1 and B7-2.
==About this Structure==
==About this Structure==
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1NCN is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Homo_sapiens Homo sapiens]. Full crystallographic information is available from [http://ispc.weizmann.ac.il/oca-bin/ocashort?id=1NCN OCA].
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1NCN is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Homo_sapiens Homo sapiens]. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1NCN OCA].
==Reference==
==Reference==
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[[Category: Homo sapiens]]
[[Category: Homo sapiens]]
[[Category: Single protein]]
[[Category: Single protein]]
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[[Category: Almo, S.C.]]
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[[Category: Almo, S C.]]
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[[Category: Nathenson, S.G.]]
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[[Category: Nathenson, S G.]]
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[[Category: Schwartz, J.D.]]
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[[Category: Schwartz, J D.]]
[[Category: Zhang, X.]]
[[Category: Zhang, X.]]
[[Category: beta strands]]
[[Category: beta strands]]
[[Category: ig v]]
[[Category: ig v]]
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''Page seeded by [http://ispc.weizmann.ac.il/oca OCA ] on Mon Nov 12 18:20:25 2007''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 14:04:39 2008''

Revision as of 12:04, 21 February 2008


1ncn, resolution 2.70Å

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the receptor-binding domain of human B7-2

Overview

B7-1 and B7-2 are homologous costimulatory ligands expressed on the surfaces of antigen-presenting cells. Their interactions with CD28/CTLA-4 receptors expressed on T cell surfaces are crucial for the proper regulation of T cell activity. B7-1 and B7-2 display distinct roles in immune regulation, although they are usually considered to have redundant functions. Here, we report the crystal structure of the receptor-binding (Ig V-type) domain of human B7-2 at 2.7-A resolution. Structures of unliganded and liganded B7-1 and B7-2 suggest a physical-chemical basis for the observed functional similarities and differences between these two costimulatory ligands. Of particular note, whereas the majority of the residues mediating B7-1 dimerization are hydrophobic, the B7-2 dimer observed in the B7-2/CTLA-4 complex displays a very hydrophilic dimer interface. These differences provide a mechanism for preventing the formation of B7-1/B7-2 heterodimers. The divergence at the putative dimer interface is also consistent with the lower tendency of B7-2 to dimerize, as shown by the monomeric state of unliganded B7-2 both in solution and crystalline form, and may result in detailed differences in signaling mechanisms associated with B7-1 and B7-2.

About this Structure

1NCN is a Single protein structure of sequence from Homo sapiens. Full crystallographic information is available from OCA.

Reference

Crystal structure of the receptor-binding domain of human B7-2: insights into organization and signaling., Zhang X, Schwartz JC, Almo SC, Nathenson SG, Proc Natl Acad Sci U S A. 2003 Mar 4;100(5):2586-91. Epub 2003 Feb 26. PMID:12606712

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