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2ogv

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==Overview==
==Overview==
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c-Fms, a member of the Platelet-derived Growth Factor (PDGF) receptor, family of receptor tyrosine kinases (RTKs), is the receptor for macrophage, colony stimulating factor (CSF-1) that regulates proliferation, differentiation and survival of cells of the mononuclear phagocyte, lineage. Abnormal expression of c-fms proto-oncogene is associated with a, significant number of human pathologies, including a variety of cancers, and rheumatoid arthritis. Accordingly, c-Fms represents an attractive, therapeutic target. To further understand the regulation of c-Fms, we, determined the 2.7 A resolution crystal structure of the cytosolic domain, of c-Fms that comprised the kinase domain and the juxtamembrane domain., The structure reveals the crucial inhibitory role of the juxtamembrane, domain (JM) that binds to a hydrophobic site immediately adjacent to the, ATP binding pocket. This interaction prevents the activation loop from, adopting an active conformation thereby locking the c-Fms kinase into an, autoinhibited state. As observed for other members of the PDGF receptor, family, namely c-Kit and Flt3, three JM-derived tyrosine residues, primarily drive the mechanism for autoinhibition in c-Fms, therefore, defining a common autoinhibitory mechanism within this family. Moreover, the structure provides an understanding of c-Fms inhibition by Gleevec as, well as providing a platform for the development of more selective, inhibitors that target the inactive conformation of c-Fms kinase.
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c-Fms, a member of the Platelet-derived Growth Factor (PDGF) receptor family of receptor tyrosine kinases (RTKs), is the receptor for macrophage colony stimulating factor (CSF-1) that regulates proliferation, differentiation and survival of cells of the mononuclear phagocyte lineage. Abnormal expression of c-fms proto-oncogene is associated with a significant number of human pathologies, including a variety of cancers and rheumatoid arthritis. Accordingly, c-Fms represents an attractive therapeutic target. To further understand the regulation of c-Fms, we determined the 2.7 A resolution crystal structure of the cytosolic domain of c-Fms that comprised the kinase domain and the juxtamembrane domain. The structure reveals the crucial inhibitory role of the juxtamembrane domain (JM) that binds to a hydrophobic site immediately adjacent to the ATP binding pocket. This interaction prevents the activation loop from adopting an active conformation thereby locking the c-Fms kinase into an autoinhibited state. As observed for other members of the PDGF receptor family, namely c-Kit and Flt3, three JM-derived tyrosine residues primarily drive the mechanism for autoinhibition in c-Fms, therefore defining a common autoinhibitory mechanism within this family. Moreover the structure provides an understanding of c-Fms inhibition by Gleevec as well as providing a platform for the development of more selective inhibitors that target the inactive conformation of c-Fms kinase.
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==Disease==
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Known diseases associated with this structure: Myeloid malignancy, predisposition to OMIM:[[http://www.ncbi.nlm.nih.gov/entrez/dispomim.cgi?id=164770 164770]]
==About this Structure==
==About this Structure==
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[[Category: Single protein]]
[[Category: Single protein]]
[[Category: Bamert, R.]]
[[Category: Bamert, R.]]
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[[Category: Broughton, S.E.]]
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[[Category: Broughton, S E.]]
[[Category: Fantino, E.]]
[[Category: Fantino, E.]]
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[[Category: Lucet, I.S.]]
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[[Category: Lucet, I S.]]
[[Category: Patel, O.]]
[[Category: Patel, O.]]
[[Category: Rossjohn, J.]]
[[Category: Rossjohn, J.]]
[[Category: Walter, M.]]
[[Category: Walter, M.]]
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[[Category: Wilks, A.F.]]
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[[Category: Wilks, A F.]]
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[[Category: Williams, N.K.]]
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[[Category: Williams, N K.]]
[[Category: macrophage colony stimulating factor receptor]]
[[Category: macrophage colony stimulating factor receptor]]
[[Category: receptor tyrosine kinase]]
[[Category: receptor tyrosine kinase]]
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Wed Jan 23 15:25:07 2008''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 18:18:19 2008''

Revision as of 16:18, 21 February 2008


2ogv, resolution 2.70Å

Drag the structure with the mouse to rotate

Crystal Structure of the Autoinhibited Human c-Fms Kinase Domain

Contents

Overview

c-Fms, a member of the Platelet-derived Growth Factor (PDGF) receptor family of receptor tyrosine kinases (RTKs), is the receptor for macrophage colony stimulating factor (CSF-1) that regulates proliferation, differentiation and survival of cells of the mononuclear phagocyte lineage. Abnormal expression of c-fms proto-oncogene is associated with a significant number of human pathologies, including a variety of cancers and rheumatoid arthritis. Accordingly, c-Fms represents an attractive therapeutic target. To further understand the regulation of c-Fms, we determined the 2.7 A resolution crystal structure of the cytosolic domain of c-Fms that comprised the kinase domain and the juxtamembrane domain. The structure reveals the crucial inhibitory role of the juxtamembrane domain (JM) that binds to a hydrophobic site immediately adjacent to the ATP binding pocket. This interaction prevents the activation loop from adopting an active conformation thereby locking the c-Fms kinase into an autoinhibited state. As observed for other members of the PDGF receptor family, namely c-Kit and Flt3, three JM-derived tyrosine residues primarily drive the mechanism for autoinhibition in c-Fms, therefore defining a common autoinhibitory mechanism within this family. Moreover the structure provides an understanding of c-Fms inhibition by Gleevec as well as providing a platform for the development of more selective inhibitors that target the inactive conformation of c-Fms kinase.

Disease

Known diseases associated with this structure: Myeloid malignancy, predisposition to OMIM:[164770]

About this Structure

2OGV is a Single protein structure of sequence from Homo sapiens. Active as Receptor protein-tyrosine kinase, with EC number 2.7.10.1 Full crystallographic information is available from OCA.

Reference

The 2.7 A crystal structure of the autoinhibited human c-Fms kinase domain., Walter M, Lucet IS, Patel O, Broughton SE, Bamert R, Williams NK, Fantino E, Wilks AF, Rossjohn J, J Mol Biol. 2007 Mar 30;367(3):839-47. Epub 2007 Jan 20. PMID:17292918

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