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1e7q

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[[Image:1e7q.jpg|left|200px]]<br /><applet load="1e7q" size="350" color="white" frame="true" align="right" spinBox="true"
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[[Image:1e7q.jpg|left|200px]]
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caption="1e7q, resolution 1.6&Aring;" />
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'''GDP 4-KETO-6-DEOXY-D-MANNOSE EPIMERASE REDUCTASE S107A'''<br />
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{{Structure
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|PDB= 1e7q |SIZE=350|CAPTION= <scene name='initialview01'>1e7q</scene>, resolution 1.6&Aring;
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|SITE= <scene name='pdbsite=AC1:Nap+Binding+Site,+Residue+Ala107+Is+Mutat+The+Native+Bei+...'>AC1</scene> and <scene name='pdbsite=AC2:Uvw+Binding+Site+For+Chain+A'>AC2</scene>
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|LIGAND= <scene name='pdbligand=SO4:SULFATE+ION'>SO4</scene>, <scene name='pdbligand=NAP:NADP+NICOTINAMIDE-ADENINE-DINUCLEOTIDE+PHOSPHATE'>NAP</scene>, <scene name='pdbligand=UVW:ACETYLPHOSPHATE'>UVW</scene> and <scene name='pdbligand=TRS:2-AMINO-2-HYDROXYMETHYL-PROPANE-1,3-DIOL'>TRS</scene>
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|ACTIVITY=
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|GENE=
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}}
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'''GDP 4-KETO-6-DEOXY-D-MANNOSE EPIMERASE REDUCTASE S107A'''
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==Overview==
==Overview==
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==About this Structure==
==About this Structure==
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1E7Q is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Escherichia_coli Escherichia coli] with <scene name='pdbligand=SO4:'>SO4</scene>, <scene name='pdbligand=NAP:'>NAP</scene>, <scene name='pdbligand=UVW:'>UVW</scene> and <scene name='pdbligand=TRS:'>TRS</scene> as [http://en.wikipedia.org/wiki/ligands ligands]. Known structural/functional Sites: <scene name='pdbsite=AC1:Nap+Binding+Site,+Residue+Ala107+Is+Mutat+The+Native+Bei+...'>AC1</scene> and <scene name='pdbsite=AC2:Uvw+Binding+Site+For+Chain+A'>AC2</scene>. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1E7Q OCA].
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1E7Q is a [[Single protein]] structure of sequence from [http://en.wikipedia.org/wiki/Escherichia_coli Escherichia coli]. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1E7Q OCA].
==Reference==
==Reference==
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Probing the catalytic mechanism of GDP-4-keto-6-deoxy-d-mannose Epimerase/Reductase by kinetic and crystallographic characterization of site-specific mutants., Rosano C, Bisso A, Izzo G, Tonetti M, Sturla L, De Flora A, Bolognesi M, J Mol Biol. 2000 Oct 13;303(1):77-91. PMID:[http://ispc.weizmann.ac.il//pmbin/getpm?pmid=11021971 11021971]
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Probing the catalytic mechanism of GDP-4-keto-6-deoxy-d-mannose Epimerase/Reductase by kinetic and crystallographic characterization of site-specific mutants., Rosano C, Bisso A, Izzo G, Tonetti M, Sturla L, De Flora A, Bolognesi M, J Mol Biol. 2000 Oct 13;303(1):77-91. PMID:[http://www.ncbi.nlm.nih.gov/pubmed/11021971 11021971]
[[Category: Escherichia coli]]
[[Category: Escherichia coli]]
[[Category: Single protein]]
[[Category: Single protein]]
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[[Category: sdr]]
[[Category: sdr]]
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 12:24:49 2008''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Mar 20 10:51:14 2008''

Revision as of 08:51, 20 March 2008


PDB ID 1e7q

Drag the structure with the mouse to rotate
, resolution 1.6Å
Sites: and
Ligands: , , and
Coordinates: save as pdb, mmCIF, xml



GDP 4-KETO-6-DEOXY-D-MANNOSE EPIMERASE REDUCTASE S107A


Overview

GDP-4-keto-6-deoxy-d-mannose epimerase/reductase is a bifunctional enzyme responsible for the last step in the biosynthesis of GDP-l-fucose, the substrate of fucosyl transferases. Several cell-surface antigens, including the leukocyte Lewis system and cell-surface antigens in pathogenic bacteria, depend on the availability of GDP-l-fucose for their expression. Therefore, the enzyme is a potential target for therapy in pathological states depending on selectin-mediated cell-to-cell interactions. Previous crystallographic investigations have shown that GDP-4-keto-6-deoxy-d-mannose epimerase/reductase belongs to the short-chain dehydrogenase/reductase protein homology family. The enzyme active-site region is at the interface of an N-terminal NADPH-binding domain and a C-terminal domain, held to bind the substrate. The design, expression and functional characterization of seven site-specific mutant forms of GDP-4-keto-6-deoxy-d-mannose epimerase/reductase are reported here. In parallel, the crystal structures of the native holoenzyme and of three mutants (Ser107Ala, Tyr136Glu and Lys140Arg) have been investigated and refined at 1. 45-1.60 A resolution, based on synchrotron data (R-factors range between 12.6 % and 13.9 %). The refined protein models show that besides the active-site residues Ser107, Tyr136 and Lys140, whose mutations impair the overall enzymatic activity and may affect the coenzyme binding mode, side-chains capable of proton exchange, located around the expected substrate (GDP-4-keto-6-deoxy-d-mannose) binding pocket, are selectively required during the epimerization and reduction steps. Among these, Cys109 and His179 may play a primary role in proton exchange between the enzyme and the epimerization catalytic intermediates. Finally, the additional role of mutated active-site residues involved in substrate recognition and in enzyme stability has been analyzed.

About this Structure

1E7Q is a Single protein structure of sequence from Escherichia coli. Full crystallographic information is available from OCA.

Reference

Probing the catalytic mechanism of GDP-4-keto-6-deoxy-d-mannose Epimerase/Reductase by kinetic and crystallographic characterization of site-specific mutants., Rosano C, Bisso A, Izzo G, Tonetti M, Sturla L, De Flora A, Bolognesi M, J Mol Biol. 2000 Oct 13;303(1):77-91. PMID:11021971

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