1nu5

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[[Image:1nu5.jpg|left|200px]]<br /><applet load="1nu5" size="350" color="white" frame="true" align="right" spinBox="true"
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[[Image:1nu5.jpg|left|200px]]
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caption="1nu5, resolution 1.95&Aring;" />
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'''Crystal structure of Pseudomonas sp. P51 Chloromuconate lactonizing enzyme'''<br />
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{{Structure
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|PDB= 1nu5 |SIZE=350|CAPTION= <scene name='initialview01'>1nu5</scene>, resolution 1.95&Aring;
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|SITE=
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|LIGAND= <scene name='pdbligand=MN:MANGANESE (II) ION'>MN</scene>
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|ACTIVITY= [http://en.wikipedia.org/wiki/Chloromuconate_cycloisomerase Chloromuconate cycloisomerase], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=5.5.1.7 5.5.1.7]
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|GENE=
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}}
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'''Crystal structure of Pseudomonas sp. P51 Chloromuconate lactonizing enzyme'''
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==Overview==
==Overview==
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==About this Structure==
==About this Structure==
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1NU5 is a [http://en.wikipedia.org/wiki/Single_protein Single protein] structure of sequence from [http://en.wikipedia.org/wiki/Pseudomonas_sp. Pseudomonas sp.] with <scene name='pdbligand=MN:'>MN</scene> as [http://en.wikipedia.org/wiki/ligand ligand]. Active as [http://en.wikipedia.org/wiki/Chloromuconate_cycloisomerase Chloromuconate cycloisomerase], with EC number [http://www.brenda-enzymes.info/php/result_flat.php4?ecno=5.5.1.7 5.5.1.7] Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1NU5 OCA].
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1NU5 is a [[Single protein]] structure of sequence from [http://en.wikipedia.org/wiki/Pseudomonas_sp. Pseudomonas sp.]. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=1NU5 OCA].
==Reference==
==Reference==
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The structure of Pseudomonas P51 Cl-muconate lactonizing enzyme: co-evolution of structure and dynamics with the dehalogenation function., Kajander T, Lehtio L, Schlomann M, Goldman A, Protein Sci. 2003 Sep;12(9):1855-64. PMID:[http://ispc.weizmann.ac.il//pmbin/getpm?pmid=12930985 12930985]
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The structure of Pseudomonas P51 Cl-muconate lactonizing enzyme: co-evolution of structure and dynamics with the dehalogenation function., Kajander T, Lehtio L, Schlomann M, Goldman A, Protein Sci. 2003 Sep;12(9):1855-64. PMID:[http://www.ncbi.nlm.nih.gov/pubmed/12930985 12930985]
[[Category: Chloromuconate cycloisomerase]]
[[Category: Chloromuconate cycloisomerase]]
[[Category: Pseudomonas sp.]]
[[Category: Pseudomonas sp.]]
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[[Category: muconate]]
[[Category: muconate]]
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Feb 21 14:10:04 2008''
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''Page seeded by [http://oca.weizmann.ac.il/oca OCA ] on Thu Mar 20 13:01:43 2008''

Revision as of 11:01, 20 March 2008


PDB ID 1nu5

Drag the structure with the mouse to rotate
, resolution 1.95Å
Ligands:
Activity: Chloromuconate cycloisomerase, with EC number 5.5.1.7
Coordinates: save as pdb, mmCIF, xml



Crystal structure of Pseudomonas sp. P51 Chloromuconate lactonizing enzyme


Overview

Bacterial muconate lactonizing enzymes (MLEs) catalyze the conversion of cis,cis-muconate as a part of the beta-ketoadipate pathway, and some MLEs are also able to dehalogenate chlorinated muconates (Cl-MLEs). The basis for the Cl-MLEs dehalogenating activity is still unclear. To further elucidate the differences between MLEs and Cl-MLEs, we have solved the structure of Pseudomonas P51 Cl-MLE at 1.95 A resolution. Comparison of Pseudomonas MLE and Cl-MLE structures reveals the presence of a large cavity in the Cl-MLEs. The cavity may be related to conformational changes on substrate binding in Cl-MLEs, at Gly52. Site-directed mutagenesis on Pseudomonas MLE core positions to the equivalent Cl-MLE residues showed that the variant Thr52Gly was rather inactive, whereas the Thr52Gly-Phe103Ser variant had regained part of the activity. These residues form a hydrogen bond in the Cl-MLEs. The Cl-MLE structure, as a result of the Thr-to-Gly change, is more flexible than MLE: As a mobile loop closes over the active site, a conformational change at Gly52 is observed in Cl-MLEs. The loose packing and structural motions in Cl-MLE may be required for the rotation of the lactone ring in the active site necessary for the dehalogenating activity of Cl-MLEs. Furthermore, we also suggest that differences in the active site mobile loop sequence between MLEs and Cl-MLEs result in lower active site polarity in Cl-MLEs, possibly affecting catalysis. These changes could result in slower product release from Cl-MLEs and make it a better enzyme for dehalogenation of substrate.

About this Structure

1NU5 is a Single protein structure of sequence from Pseudomonas sp.. Full crystallographic information is available from OCA.

Reference

The structure of Pseudomonas P51 Cl-muconate lactonizing enzyme: co-evolution of structure and dynamics with the dehalogenation function., Kajander T, Lehtio L, Schlomann M, Goldman A, Protein Sci. 2003 Sep;12(9):1855-64. PMID:12930985

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