3zf8

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{{STRUCTURE_3zf8| PDB=3zf8 | SCENE= }}
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==Crystal structure of Saccharomyces cerevisiae Mnn9 in complex with GDP and Mn.==
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===Crystal structure of Saccharomyces cerevisiae Mnn9 in complex with GDP and Mn.===
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<StructureSection load='3zf8' size='340' side='right' caption='[[3zf8]], [[Resolution|resolution]] 1.98&Aring;' scene=''>
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{{ABSTRACT_PUBMED_24026536}}
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== Structural highlights ==
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<table><tr><td colspan='2'>[[3zf8]] is a 1 chain structure with sequence from [http://en.wikipedia.org/wiki/Saccharomyces_cerevisiae_s288c Saccharomyces cerevisiae s288c]. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=3ZF8 OCA]. For a <b>guided tour on the structure components</b> use [http://oca.weizmann.ac.il/oca-docs/fgij/fg.htm?mol=3ZF8 FirstGlance]. <br>
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==Function==
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</td></tr><tr id='ligand'><td class="sblockLbl"><b>[[Ligand|Ligands:]]</b></td><td class="sblockDat"><scene name='pdbligand=GDP:GUANOSINE-5-DIPHOSPHATE'>GDP</scene>, <scene name='pdbligand=MN:MANGANESE+(II)+ION'>MN</scene></td></tr>
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<tr id='resources'><td class="sblockLbl"><b>Resources:</b></td><td class="sblockDat"><span class='plainlinks'>[http://oca.weizmann.ac.il/oca-docs/fgij/fg.htm?mol=3zf8 FirstGlance], [http://oca.weizmann.ac.il/oca-bin/ocaids?id=3zf8 OCA], [http://www.rcsb.org/pdb/explore.do?structureId=3zf8 RCSB], [http://www.ebi.ac.uk/pdbsum/3zf8 PDBsum]</span></td></tr>
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</table>
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== Function ==
[[http://www.uniprot.org/uniprot/MNN9_YEAST MNN9_YEAST]] The M-Pol I and M-Pol II complexes possess alpha-1,6-mannosyltransferase activity and are probably involved in the elongation of the mannan backbone of N-linked glycans on cell wall and periplasmic proteins. May also provide alpha-1,2-mannosyltransferase activity to the M-Pol I complex.<ref>PMID:12235155</ref>
[[http://www.uniprot.org/uniprot/MNN9_YEAST MNN9_YEAST]] The M-Pol I and M-Pol II complexes possess alpha-1,6-mannosyltransferase activity and are probably involved in the elongation of the mannan backbone of N-linked glycans on cell wall and periplasmic proteins. May also provide alpha-1,2-mannosyltransferase activity to the M-Pol I complex.<ref>PMID:12235155</ref>
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<div style="background-color:#fffaf0;">
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== Publication Abstract from PubMed ==
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The fungal cell possesses an essential carbohydrate cell wall. The outer layer, mannan, is formed by mannoproteins carrying highly mannosylated O- and N-linked glycans. Yeast mannan biosynthesis is initiated by a Golgi-located complex (M-Pol I) of two GT-62 mannosyltransferases, Mnn9p and Van1p, that are conserved in fungal pathogens. Saccharomyces cerevisiae and Candida albicans mnn9 knockouts show an aberrant cell wall and increased antibiotic sensitivity, suggesting the enzyme is a potential drug target. Here, we present the structure of ScMnn9 in complex with GDP and Mn(2+), defining the fold and catalytic machinery of the GT-62 family. Compared with distantly related GT-78/GT-15 enzymes, ScMnn9 carries an unusual extension. Using a novel enzyme assay and site-directed mutagenesis, we identify conserved amino acids essential for ScMnn9 'priming' alpha-1,6-mannosyltransferase activity. Strikingly, both the presence of the ScMnn9 protein and its product, but not ScMnn9 catalytic activity, are required to activate subsequent ScVan1 processive alpha-1,6-mannosyltransferase activity in the M-Pol I complex. These results reveal the molecular basis of mannan synthesis and will aid development of inhibitors targeting this process.
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==About this Structure==
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Yeast Mnn9 is both a priming glycosyltransferase and an allosteric activator of mannan biosynthesis.,Striebeck A, Robinson DA, Schuttelkopf AW, van Aalten DM Open Biol. 2013 Sep 11;3(9):130022. doi: 10.1098/rsob.130022. PMID:24026536<ref>PMID:24026536</ref>
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[[3zf8]] is a 1 chain structure with sequence from [http://en.wikipedia.org/wiki/Saccharomyces_cerevisiae_s288c Saccharomyces cerevisiae s288c]. Full crystallographic information is available from [http://oca.weizmann.ac.il/oca-bin/ocashort?id=3ZF8 OCA].
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==Reference==
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From MEDLINE&reg;/PubMed&reg;, a database of the U.S. National Library of Medicine.<br>
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<ref group="xtra">PMID:024026536</ref><references group="xtra"/><references/>
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</div>
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== References ==
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<references/>
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__TOC__
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</StructureSection>
[[Category: Saccharomyces cerevisiae s288c]]
[[Category: Saccharomyces cerevisiae s288c]]
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[[Category: Aalten, D M.F van.]]
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[[Category: Aalten, D M.F van]]
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[[Category: Schuettelkopf, A W.]]
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[[Category: Schuettelkopf, A W]]
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[[Category: Striebeck, A.]]
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[[Category: Striebeck, A]]
[[Category: Transferase]]
[[Category: Transferase]]

Revision as of 15:39, 24 December 2014

Crystal structure of Saccharomyces cerevisiae Mnn9 in complex with GDP and Mn.

3zf8, resolution 1.98Å

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