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Glutathione Reductase (GSR) reduces glutathione disulfides (molecule with two yellow spheres shown in the 3D graphic) to
Glutathione is an anti-oxidant molecule (which itself is made up of three amino acid molecules -glutamine, cystine and glycine) that keeps cells in "shape" ensuring other proteins in the cell soup don't get oxidized and keep the cell machinery running.
flavin adenine dinucleotide (flat molecule) another molecule that helps GSR to break glutathione disulfide bonds is juxtaposed up close to the glutathione disulfide molecule.
Wikipedia page on GSR